Haplogroup T-M184


Haplogroup T-M184, also known as Haplogroup T, is a human Y-chromosome DNA haplogroup. The unique-event polymorphism that defines this clade is the single-nucleotide polymorphism known as M184.
T-M184 is unusual in that it is both geographically widespread and relatively rare. T1 – the numerically dominant primary branch of T-M184 – appears to have originated in Western Asia, and spread from there into East Africa, South Asia, Europe, Egypt and adjoining regions. T1* may have expanded with the Pre-Pottery Neolithic B culture which originated in West Asia.
The earliest presence of T-M184 appears in Ain Ghazal, Jordan, bordering Asia and Africa. The individual predated the arrival of Caucaso-Iranian ancestry to the Levant. His DNA consisted of Natufian Hunter Gatherer and Anatolian Neolithic ancestry, together known as PPNB, which was the indigenous ancestry of the Levant at the time.
Subclades of T-M70 appear to have been present in Europe since the Neolithic with Neolithic Farmers from Western Asia. The moderately high frequency of T1b* chromosomes in the Lemba of southern Africa supports the hypothesis of a West Asian origin for their paternal line.

Structure

Haplogroup_K = ~45,000–50,000 years ago",
    "Region": "South or Southeast Asia"
"Descendants" "LT "~40,000–45,000 years ago",
"Region": "South Asia / Iranian Plateau", "
Descendants": L~25,000–30,000 years ago",
  "Region": "South Asia, Dravidians, Indus Valley",
 "Notes": "Possibly linked to pre-Neolithic South Asian populations
     |  "T": ~25,000–30,000 years ago", |
       "Region": "South Asia,",
        "Notes": "Later spread into the Levant, Mediterranean, and Africa"
;Subclade structure of Haplogroup T.
  • T1
  • *T1a
  • **T1a1
  • ***T1a1a
  • ****T1a1a1
  • ****T1a1a2
  • *****T1a1a2a
  • **T1a2
  • ***T1a2a
  • ***T1a2b
  • **T1a3
  • ***T1a3a
  • ***T1a3b
  • '''T2'''

Distribution

Overview

As a primary branch of haplogroup LT, the basal, undivergent haplogroup T* currently has the alternate phylogenetic name of K1b and is a sibling of haplogroup L*. It has two primary branches: T1 and T2. Most males who now belong to haplogroup T1* carry the subclade T-M70, a primary branch of T-M206.
Haplogroup T is found at exceptionally high levels amongst the Dir and Isaaq in Somaliland, Djibouti, and Ethiopia. it is also found at relatively high levels in specific populations in other parts of the world especially amongst Arabs from UAE in South Eastern Arabia T-M184 spikes at 19% on FTDNA. These include Kurru, Bauris and Lodha in South Asia; among Toubou in Chad; Somalilander clans: Isaaq and Dir, southern Egyptians and Fula in north Cameroon; ; Zoroastrians, Bakhtiaris, Assyrians and Iraqi Jews in the Middle East. T is a rather rare haplogroup, displaying a global frequency of around 1%, but nonetheless it is found at quite high frequencies in Sephardic Levites and Sephardic Israelis.
The maximal worldwide frequency for haplogroup T-M184 is 100%, amongst Dir clan males. It accounts for approximately 82.4% of ethnic Somali male lineages overall in Dire Dawa, Ethiopia. T is only 9% in Somalia . Geographically, it is found at the highest levels in the Dire Dawa area of Ethiopia, and Djibouti.
Luis et al. suggest that the presence of T on the African continent may, like R1* representatives, point to an older introduction from West Asia. The Levant rather than the Arabian Peninsula appears to have been the main route of entry, as the Egyptian and Anatolian haplotypes are considerably older in age than those found in Oman. According to the authors, haplogroup T-M184 within Africa represents the traces of a more widespread early local presence of the West Asian clade. Later expansions of populations from West Asia carrying the E-M215, E-V38, G and J NRY lineages may have overwhelmed the T-M184 clade-bearers in certain localities.
In the Caucasus and Anatolia it makes up to 4% of the population in southeast and northwest Caucasus as well as in southeast and western Anatolia, peaking up to 20% in Armenians from Sasun. In Middle East it makes up to 4% of the population around the Zagros Mountains and the Persian Gulf as well as around the Taurus Mountains and the Levant basin, peaking up to 10% in Zoroastrians from Kerman, Bakhtiaris, Assyrians, Abudhabians, Armenians from Historical Southwestern Armenia and Druzes from Galilee. In Eastern Africa, it makes up to 4% of the population on Upper Egypt peaking up to 10% in Luxor.
Haplogroup T is uncommon in Europe, except in Southern Europe and adjoining areas. According to Mendez et al., "the occurrence in Europe of lineages belonging to both T1a1 and T1a2 subclades probably reflects multiple episodes of gene flow. T1a1* haplogroups in Europe likely reflect older gene flow". It makes up to 4% of the population in Central Italy, Western Sicily, Northwest Corsica, Northwestern Iberian Peninsula, Western Andalucia, Western Alps, Eastern Crete, and Macedonia, frequencies up to 10% in Ibiza, Miranda de I Douro, Eastern Oviedo, Cádiz, Badajoz, Balagna, Norma and Ragusa, and peaking at 20% in Sciacca, L'Aquila and some German southern regions. T-M184 was found in 1.7% of a pool of six samples of males from southwestern Russia, but it was completely absent from a pool of eight samples totalling 637 individuals from the northern half of European Russia. The Russians from the southwest were from the following cities: Roslavl, Livny, Pristen, Repyevka, and Belgorod; and Kuban Cossacks from the Republic of Adygea.

T1 (T*)

T1 is the most common descent of T-M184 haplogroup, being the lineage of more than 95% of all Eurasian T-M184 members. One of their descent lineages is found in high frequencies among northern Somali clans. However, it appears to have originated somewhere around the Eastern Mediterranean Basin, perhaps somewhere between Palestine to the Jordan Valley.
The basal T1* subclade appears to have spread to northeastern Anatolia, from the Levant and Mesopotamia at least, with the Pre-Pottery Neolithic B culture. Although it is rare in modern populations, T1* has been found in a Berber individual from Tunisia, a male in Syria, and one sequence among ethnic Macedonians in Macedonia.

T1a (M70)

Mendez et al. points to an ancient presence for T1a-M70 in Europe may reflect early exiles between the ancient lands of Israel and Babylonia and Assyria. The subclade probably arrived with the very first farmers.

T1a1*

The Pityusans of the Pityusic Islands – have been found by three different studies to possess T1a1 at relatively high levels of 6.7–16.7%. Tomàs et al. found three cases amongst a sample of 45. Zalloua et al. found nine examples that were L454+ from a sample of 54. Rodriguez et al. found seven cases of L454+ in a sample of 96.
The Pontic Greeks of Anatolia are also reported to possess T1a1. In 2009, a male with the surname Metaxopoulos and a Pontic Greek background was reported to be T-L162 – according to the Y-Chromosome Genome Comparison Project administered by Adriano Squecco. Greeks from the Fatsa reportedly migrated in antiquity from Sinope, which was itself colonised by Ionians. Another ancient Ionian colony in north-west Anatolia, Lámpsakos, had onomastic links to the Pityusic Islands – Lámpsakos was originally an Ionian colony known as Pityussa.

T1a1a (L208)

This lineage, formed 14,200-11,000 BP, is the largest branch downstream T1a1-L162. The Isaaq clans and dir is T-L208, they found in Somaliland, Djibouti, Somalia and Somali Ethiopia.

T1a1a1a1b1a1* (T-Y3782*)

One Sardinian male from a sample of 187 – a resident of the Province of Cagliari – has been found to have T-Y3782, also known T1a1a1a1b1a1.

T1a1a1a1b1a1a (T-Y3836)

This lineage is mostly found among individuals from the Iberian Peninsula, where the subclade also has its highest diversity. Two subclades can be clearly discriminated. The first, found mainly in post-colonial Puerto Rico, with DYS391=10 and the second, found mainly in Panamá where their Iberian descendants could have the entrance point to America, with DYS439=12.
Some members of Y3836 are found among different communities of the Sephardic diaspora but they are found to be extremely rare in the total percentage of some of these communities as seen in Nogueiro et al. This probably could mean that these members could be integrated by these communities through the contact with other native Iberian populations as seen in Monteiro et al. where this lineage was found among native Astur-Leonese speakers.
PopulationLanguageLocationMembers/Sample sizePercentageSourceNotes
PanamaniansPanamian Castilian Los Santos Province1/303.3%
ColombiansColombian Castilian Caldas2/752.7%YHRDMestizo individuals
PanamaniansPanamian Castilian Panama Province1/432.3%
Northwest ArgentiniansArgentinian Castilian Mountainous region of Jujuy1/502%YHRDAdmixed population
Puerto RicansPuerto Rican Castilian Southeast Puerto Rico2/1101.8%
Northeastern Portuguese JewsJudaeo-Portuguese Bragança, Argozelo, Carção, Mogadouro, and Vilarinho dos Galegos1/571.8%
Native Mirandese speakersMirandese Astur-Leonese Miranda de l Douro1/581.7%
DominicansDominican Castilian Dominican Republic4/2611.5%
PanamaniansPanamian Castilian Chiriquí Province1/921.1%
MecklenburgersEast Low Saxon Rostock2/2001%
MestizosColombian Castilian Bogotá2/1951%YHRD
MestizosColombian Castilian Valle del Cauca1/1031%YHRD
MestizosEcuadorian Castilian Quito1/1021%
VenezuelansVenezuelan Castilian Maracaibo1/1110.9%
VenezuelansVenezuelan Castilian Central Region1/1150.9%
EuropeansBrazilian Portuguese São Paulo1/1200.8YHRDEuropean descents
EcuadoriansEcuadorian Castilian Quito1/1200.8%
ColombiansColombian Castilian Antioquia6/7770.7%
MexicansMexican Castilian Mérida1/1590.6%YHRDMestizo individuals
Eastern AndalusiansAndalusian Alhama de Granada, Baza, Huéscar, Loja, Montefrío and Órgiva1/1800.6%
ColombiansColombian Castilian Santander1/1930.5%YHRDMestizo individuals
ChileansChilean Castilian Concepción1/1980.5%YHRD
CatalansNot reportedMetropolitan area of Barcelona1/2240.5%
MexicansMexican Spanish Guadalajara1/2460.4%YHRDMestizo individuals
EuropeansBrazilian Portuguese Rio Grande do Sul1/2550.4%

T2 (PH110)

This lineage could have arrived in the Levant through the PPNB expansion from northeastern Anatolia.
A 2014 study found T-PH110 in one ethnic Bhutanese male, out of a sample of 21, possibly implying a rate of 4.8% in Bhutan. Also have been found in a German individual and another two from Caucasus. The Bhutanese and the German haplotypes seems to cluster together.

Possible cases from older research

PopulationLanguageLocationMembers/Sample sizePercentageSourceNotes
AltaiansAltai Kurmach-Baygol2/1118.2%K*
AltaiansAltai Turochak2/1910.5%K
LeonesesAstur-Leonese Leon1/137.7%K
Ossetian IronsIron South Ossetia1/214.8%No further details available.
CordobesesAndalusian Córdoba1/273.7%No further details available.
LeonesesAstur-Leonese Leon2/603.3%No further details available.
TharusTharu Morang1/372.7%K
CherkessiansBesleney Circassia2/1261.6%No further details are available.
BizkaiansBizkaiera Bizkaia1/721.4%No further details are available.
EuropeansEnglish Australia1/10780.09%No further details are available.

Modern geographical distribution

Europe

With K-M9+, unconfirmed but probable T-M70+: 14% of Russians in Yaroslavl, 12.5% of Italians in Matera, 10.3% of Italians in Avezzano, 10% of Tyroleans in Nonstal, 10% of Italians in Pescara, 8.7% of Italians in Benevento, 7.8% of Italians in South Latium, 7.4% of Italians in Paola, 7.3% of Italians in Central-South Italy, 7.1% of Serbs in Serbia, 4.7% of Aromanians in Romania, 3.7% of Italians in Biella, 3.7% of Andalusians in Córdoba, 3.3% of Leoneses in León, 3.2% of Italians in Postua, 3.2% of Italians in Cavaglià, 3.1% of Calabrians in Reggio Calabria, 2.8% of Russians in Ryazan Oblast, 2.8% of Italians in South Apulia, 2.7% of Calabrians in Cosenza, 2.6% of Serbs in Belgrade, 2.5% of Russians in Pskov, 2.4% of Russians in Kaluga, 2.2% of Transylvanians in Miercurea Ciuc, 2.2% of Italians in Trino Vercellese, 1.9% of Italians in Brescia, 1.9% of Romanians in Romania, 1.7% of Serbs and Montenegrins in Serbia and Montenegro, 1.7% of Italians in Marche, 1.7% of Calabrians in Catanzaro, 1.6% of Greeks in Northern Greece, 1.3% of Swiss Germans in Zürich Area, 1.3% of Italians in South Tuscany and North Latium, 1.1% of Dutch in Leiden, 0.5% of Serbs in Novi Sad, 0.5% of Polish in Podlasie
Other parts that have been found to contain a significant proportion of haplogroup T-M184 individuals include Trentino, Mariña Lucense, Heraklion, Roslavl, Ourense, Livny, Biella, Entre Douro, Porto, Urbino, Iberian Peninsula, Blekinge/Kristianstad, Belarus, Modena, Provence-Alpes-Côte d'Azur, Pristen, Cáceres, Brac, Satakunta, Western Croatia, Ukrainia, Greifswald, Moldavians in Sofia, Uppsala, Lublin, Pias in Beja, Macedonian Greeks, Nea Nikomedeia, Sesklo/Dimini, Lerna/Franchthi, Açores, Viana do Castelo, Toulouse, Belgorod, Sardinia. According to data from commercial testing, 3.9% of Italian males belonging to this haplogroup. Approximately 3% of Sephardi Jews and 2% of Ashkenazi Jews belong to haplogroup T.

Middle East and Caucasus

Haplogroup T has some significant frequencies in southeast and eastern Anatolia, the Zagros Mountains and both sides of the Persian Gulf.
PopulationLanguageLocationMembers/Sample sizePercentageSourceNotes
GeorgiansGeorgian Khashuri1/333.3%
Priest ZoroastriansPersianShiraz, Tehran and Yazd2/825%Not specified if Herbad or Mobad
Iraqi JewsJudeo-Iraqi Arabic Iraq7/3221.9%12.5% T1a1a1a1a1a1-P77 and 9.4% T1a3-Y11151
Armenian SasuntzisWestern Armenian dialect, Kurmanji and Dimli languagesSasun21/10420.2%T1a1 and T1a2 subclades
GeorgiansGeorgian Sighnaghi and Gurjaani2/1020%
GeorgiansGeorgian Kharagauli1/520%
KumyksKumyk Daghestani lowlands2/1020%Reported as K* but according to Karafet16 and Yunusbayev12 only T fits.
Kurdish JewsJudeo-Aramaic Kurdistan19/9919.2%
Kurdish JewsJudeo-Aramaic Kurdistan9/5018%10% T1a1a1a1a1a1-P77 and 8% T1a1-L162
DruzesPalestinian Arabic Galilee7/4017.5%
AssyriansAramaic refugees in Armenia16/10615.1%Reported as K*. Their homeland in the areas around Urmia.
AssyriansAramaic Unknown4/2814.3%
GeorgiansGeorgian Dusheti1/714.3%
Iranian JewsJudeo-Iranian Iran3/2213.6%4.5% T1a1a1a1a1a1-P77 and 9.1% T1a3-Y11151
ZoroastriansPersianKerman5/3713.5%
Iraqi JewsJudeo-Iraqi Arabic Iraq13/9913.1%
BakhtiarisBakhtiari Izeh13/10312.6%
Mountain JewsJudeo-Tat Derbentsky District2/1711.8%All belong to T1a1a1a1a1a1-P77
ArmeniansWestern Armenian dialectHistorical Southwestern Armenia11/9611.5%
EmiratisGulf Arabic Abu Dhabi21/19111%
AssyriansAssyrian West Azerbaijan Province4/3910.3%
Iranian JewsJudeo-Iranian Iran5/4910.2%
Persian MuslimsPersianShiraz5/519.8%
Persian MuslimsPersianKerman6/669.1%
IraqisIraqi Arabic Al-Qadisiyah6/698.7%
ArmeniansArmenianArmenia35/4138.5%
KurdsSorani Kurdestan5/598.5%
Omani ArabsOmani Arabic Oman10/1218.3%
KurdsSorani Kurdestan2/258%
AzerisAzeri West Azerbaijan Province5/637.9%
MazanderanisMazanderan Mazandaran1/137.7%
CypriotsCypriot GreekCyprus3/417.3%
IraqisIraqi Arabic Iraq10/1397.2%
KuwaitisGulf Arabic Kuwait3/427.1%
IraqisIraqi Arabic Iraq3/437%
ArabsLevantine ArabicIsrael and Palestine10/1437%
PersiansFarsi Fars3/446.8%
Christian ArabsLevantine ArabicIsrael and Palestine3/446.8%
Western ArmeniansArmenianEastern Turkey6/906.7%
PersiansFarsi Yazd3/466.5%
ArmeniansArmenianGardman6/966.3%
YezidisKurmanji refugees in Armenia12/1966.1%Reported as K*. Their homeland in the areas around Laliş.
Muslim ArabsLevantine ArabicIsrael and Palestine7/1195.9%
Zahedan, Baluchestan, Iran6/1035.8%
Northern ArmeniansArmenianNorthern Armenia, southern Georgia and northwestern Azerbaijan 10/1895.3%
ArmeniansArmenianTehran2/385.3%
Eastern ArmeniansArmenianKarabakh11/2155.1%
PersiansFarsi Khorasan3/595.1%
Saudi ArabiansArabic dialects Saudi Arabia8/1575.1%
ArmeniansArmenianSyunik7/1405%
EmiratisGulf Arabic United Arab Emirates8/1644.9%
Lebanese MuslimsLebanese Arabic Lebanon28/5684.9%
CypriotsCypriot GreekLemesos6/1264.8%
KumyksKumyk Khasavyurtovsky District1/214.8%
AvarsAvar southeastern Dagestan2/424.8%
KurdsKurmanji Anatolia12/2514.8%
KurdsKurdish dialects Kurdistan6/1264.8%
AnizesGulf Arabic Kuwait1/214.7%
LebanesesLevantine Arabic Lebanon43/9144.7%
CypriotsCypriot GreekCyprus3/654.6%
MaronitesLebanese Arabic and Syriac Lebanon24/5184.6%
ArmeniansArmenianArarat2/444.6%
Muslim KurdsKurdish dialects Kurdistan4/954.2%
QeshmisQishmi Qeshm2/494.1%
LursLuri Lorestan2/504%
SadatsLanguages of IranDifferent cities of Iran2/504%
PersiansPersianEastern Iran3/773.9%
ArmeniansArmenianLake Van4/1033.9%
Saudi ArabiansArabic dialects Saudi Arabia4/1063.8%
Turkish CypriotsCypriot Turkish138 different villages, towns or cities from Cyprus14/3803.7%Paternal lineages originating from the traditional Turkish Cypriot settlements throughout the island
Birjand, South Khorasan, Iran1/273.7%All T1a3-Y12871
ArmeniansArmenianArarat Valley4/1103.6%
ArmeniansArmenianArmenia2/573.5%
GeorgiansGeorgian Omalo1/293.5%
IraniansLanguages of IranSouth Iran4/1173.4%
IoniansGreekPhokaia1/313.2%
BandarisBandari Bandar Abbas4/1313.1%
CypriotsCypriot GreekLarnaka2/673%
AlansKarachay-Baksan-Chegem Kabardino-Balkaria1/692.9%
JordaniansArabic dialects Jordan8/2732.9%
CypriotsCypriot GreekAmmochostos3/1222.5%
LezghinsLezgian Southern Dagestan2/812.5%
TurksTurkishTurkey13/5232.5%
PersiansPersian Esfahan1/132.4%
IraniansLanguages of IranIran7/3242.2%
Azerbaijani MuslimsAzerbaijani Uromia2/912.2%
Yemenite JewsHebrew and ArabicYemen2/942.1%
AndisAndi western Dagestan1/492%
CypriotsCypriot GreekPaphos2/1051.9%
CypriotsCypriot GreekNicosia3/1611.9%
AssyriansAssyrian Neo-Aramaic Uromia and Tehran1/551.8%
AbkhaziansAbkhaz Abkhazia1/581.7%
KuwaitisGulf Arabic Kuwait2/1171.7%
Greek OrthodoxKoine GreekLebanon2/1161.7%
Mashhad, Razavi Khorasan, Iran2/1291.6%0.8% T1a3-Y11151
AeoliansGreekSmyrna1/681.5%
GeorgiansGeorgian Georgia1/661.5%
TurkmensTurkmen Golestan1/681.5%
KumyksKumyk Northern Dagestan1/731.4%
Kuban NogaysNogai north of Sea of Azov around Prymorsk1/871.2%
Ossetian DigorsDigorian North Ossetia1/1270.8%
Yemeni ArabsSanaani Arabic Sana'a1/1290.8%
SyriansSyrian Arabic Syria4/5180.8%
KabardinsKabardian Kabardino-Balkaria1/1400.7%
CircassiansAdyghe Republic of Adygea1/1420.7%
AbkhaziansAbkhaz Abkhazia1/1620.6%

There are also unconfirmed reports of T-M70+ amongst 28% of Lezginians in Dagestan, 21.7% of Ossetians in Zamankul, 14% of Iranians in Isfahan, 13% of Ossetians in Zil'ga, 12.6% of Kurmanji Kurds in Eastern Turkey, 11.8% of Palestinian Arabs in Palestine, 8.3% of Iranians in Shiraz, 8.3% of Ossetians in Alagir, 8% of Kurmanji Kurds in Georgia, 7.5% of Iranians in Tehran, 7.4% of Palestinian Arabs in Israeli Village, 7% of Palestinian Arabs in Israel and Palestine, 5% of Chechens in Chechenia, 4.2% of Azerbaijanians in Azerbaijan, 4.1% of Iranians in Isfahan, 4% of Armenians in Armenia, 4% of Bedouins in Israel and 2.6% of Turks in Ankara.

Africa

Fossils excavated at the Late Neolithic site of Kelif el Boroud in Morocco, which have been radiocarbon-dated to around 3,000 BCE, have been found to belong to haplogroup T-M184.
PopulationLanguageLocationMembers/Sample sizePercentageSourceNotes
Somalis Somali Djibouti24/24100%The main sub-clans of the Dir clan in Djibouti are the Issa and Gadabuursi.
Somalis Somali Dire Dawa14/1782.4%Dir sub-clans of Dire Dawa are Issa, Gurgura and Gadabuursi.
AnteonyAntemoro old Antemoro Kingdom22/3759.5%The Anteony are the descendants of aristocrats, from whom the Antemoro king is chosen. Can be grouped into the Silamo, because they have the right to undertake the ritual slaughter of animals
Somalis and AfarsSomali and Afar Djibouti30/5456.6%Mixed sample of Somali and Afar individuals.
Somalis Somali Shilavo (woreda) 5/1050%The geographic location of this Ethiopia sample as seen in Fig.1.
Somalis Somali Somaliland4/4100%All belonging to the T1a-Y16897 subclade
AfarsAfar language Djibouti5/2025%
ToubouToubouChad31%All belonging to the T1a-PF5662 subclade
AkieAkie people Tanzania3/1323.1%Akie people have remnants of a Cushitic language
SomalisSomali Jijiga 19/8322.9%Jijiga Somalis.
Arabs from SomaliaSomali immigrants in Yemen7/3321.2%
LembaVenda and Shona South Africa6/3417.6%Exclusively belong to T1a2*. Possible recent founder effect. Low frequency of T1a2 has been observed in Bulgarian Jews and Turks but is not found in other Jewish communities. Y-str Haplotypes close to some T1a2 Armenians.
RangiRangi Language Tanzania5/3215.6%
-Somalia15/10514.3%-
IraqwIraqw language Tanzania6/4712.8%
WachaggaKichagga Dār as-Salām3/2412.5%Mixed with Rift Southern Cushites.
SomaliSomali immigrants to Norway12/10411.5%
BenchBenchBench Maji Zone14/12611.4%
KoresSNNP2/1811.1%
OromoAfaan Oromo language Oromiyaa1/911.1%
FulbeFulanorthern Cameroon3/2711.1%
GorowaGorowa language Tanzania2/1910.5%
SomaliSomali immigrants to Denmark21/20110.4%
Upper EgyptiansEgyptian ArabicLuxor Governorate3/2910.3%
KontasKonta language Konta special woreda11/10710.3%
RendilleRendille language Marsabit County3/319.7%
DatogsRendille language Tanzania3/319.7%
GewadasGewada language SNNP11/1169.5%
AntalaotraAntemoro old Antemoro Kingdom4/439.3%The Antalaotra are in charge of the magical and religious domains; they have the ability to read and write Sorabe. Can be grouped into the Silamo, because they have the right to undertake the ritual slaughter of animals
Upper EgyptiansEgyptian ArabicAswan Governorate1/119.1%
N'Djamena MixMixN'Djamena5/559.1%Marc Haber 2016All belonging to the T1a-PF5662 subclade
Upper EgyptiansEgyptian ArabicAssiut Governorate6/708.6%
KonsosKonso special woreda2/248.3%
SomaliSomali immigrants to Sweden12/1478.2%
Arabs and BerbersEgyptian Arabic and SiwiLower Egypt12/1478.2%
Upper EgyptiansEgyptian ArabicSohag Governorate4/527.7%
EgyptiansErythraic Egypt7/927.6%If the K* sample is M184+ then 8.7%
TigrayansTigrinya Tigray Region2/306.7%
DirashasDirasha Dirashe special woreda5/796.3%
CanariansCanarian SpanishTenerife11/1786.2%
KordofaniansKordofanianKurdufan4/695.8%
Upper EgyptiansEgyptian ArabicQena Governorate3/525.8%
TuaregTuareg Gorom-Gorom1/185.6%
AfarsAfar Afar Region6/1115.4%
EthiopiansEthiopian languagesEthiopia4/745.4%
MashilesMashile language SNNP7/1305.4%
GuragesGurage languages SNNP6/1185.1%
TuruNyaturu Tanzania1/205%
Moroccan JewsHaketia Israel1/205%
GedeosGedeo SNNP6/1224.9%
WairakIraqw Tanzania2/414.9%
Western LibyansLibyan Arabic Tripoli region7/1424.9%
TunisiansTunisian Arabic Sfax5/1054.8%
LibyansLibyan Arabic Tripoli area3/634.8%
KanuriKanuriCameroon1/214.8%
IraqwIraqw Tanzania2/434.7%
YemsYemsa SNNP5/1074.7%
JewsEthiopia1/224.5%
GobezeCushiticSNNP5/1134.4%
Upper EgyptiansEgyptian ArabicMinya Governorate1/234.3%
KonsosKonso language Konso special woreda4/944.3%
KembaatasEast CushiticKembata Tembaro Zone4/1023.9%
TigrayansTigrinya Eritrea1/283.6%
TigrayansTigrinya Eritrea1/313%
AmharasAmharic Ethiopia1/342.9%
HutusRwanda-Rundi Rwanda1/392.6%
Lower EgyptiansEgyptian Arabic Mansoura1/442.2%
BerbersShilha Siwa Oasis2/932.2%
MeruMeru Tanzania2/992%
ItamIbibioObong Itam 1/502%
Cape VerdeansCape Verdean Creole Windward islands São Nicolau, São Vicente, and Santo Antão2/1012%
OvimbundoUmbundu and PortugueseAngola1/531.9%
TunisiansTunisian Arabic Tunis1/541.9%
BerbersShilha Asni1/541.9%
Eastern LibyansLibyan Arabic Benghazi4/2141.9%
AlgeriansAlgerian Arabic Algeria3/1641.8%
BaribasBaatonum Benin1/571.8%T1a-M70
BokorasKaramojong Karamoja region1/591.7%
Lower EgyptiansEgyptian Arabic Cairo1/631.6%
TumbukaTumbuka northern Malawi1/611.6%
MozabitesMozabite Ghardaia1/681.5%
TunisiansTunisian Arabic South Tunisia3/2001.5%
SoussiansTunisian Arabic Sousse3/2201.4%
ChewaChewa Malawi1/921.1%
MaasaiMaasai Kinyawa (Mashuru)1/1001%YHRD
BantuNarrow Bantu Pretoria1/981%
NilotesAteker Karamoja region1/1180.8%
AndalusiansAndalusian Arabic Testour, El Alia, Gualaat-El-Andalous, Slouguia1/1320.8%Refugees from Al-Andalus following the capitulation of the Islamic kingdoms in Valencia and Granada
BantusBantuBotswana, Namibia and Zambia1/1400.7%Father and paternal grandfather belonged to the same ethnolinguistic group
BasothosSesotho Lesotho1/1810.6%
MoroccansMoroccan Arabic Casablanca metropolitan area1/1660.6%The industrial capital of Morocco where the urban growth is maintained by immigration from all parts of Morocco
KhoisansKhoisanBotswana, Namibia and Zambia1/3710.3%Father and paternal grandfather belonged to the same ethnolinguistic group

South Asia

T1a-M70 in India has been considered to be of West Eurasian origin.
PopulationLanguageLocationMembers/Sample sizePercentageSourceNotes
KurruYerukala Andhra Pradesh10/1855.6%
BaurisBengali West Bengal10/1952.6%K* is found at 6/19, if M70- but M184+, then could be 84.2%.
LodhaLodhi West Bengal2/450%
RajusTelugu Andhra Pradesh3/1915.9%
MaheliMahali West Bengal2/1315.3%
ChenchusChenchu Andhra Pradesh3/2015%K* is found at 7/20, if M70- but M184+, then could be 50%
Kare VokkalKannada Uttara Kannada4/3013.3%K* is found at 3/30, if M70- but M184+, then could be 23.3%
BanjarasLambadi Andhra Pradesh2/1811.1%
GondsGondi South Uttar Pradesh4/3810.6%
GondsGondi Madhya Pradesh10/1397.2%
Indianslanguages of IndiaSouth India18/3055.9%
MaheliMahali Jamshedpur from Jharkhand; Purulia, Midnapore & other location from West Bengal2/385.3%Two samples from different studies grouped together
ChenchusChenchu Andhra Pradesh3/614.9%Samples from Trivedi et al. and Kivisild et al.
BanjarasLambadi Andhra Pradesh2/533.8%Two samples from different studies grouped together
Indianslanguages of IndiaEast India14/3673.8%
GujaratisGujarati Gujarat1/293.4%
LodhaLodhi Midnapore & other location from West Bengal2/712.8%Three samples from different studies grouped together
SahariyasSaharia Madhya Pradesh2/732.7%
TamtasBageshwar1/342.9%
KshatriyasPithoragarh2/792.5%
AryasArya Nainital1/462.2%
LaotiansLao Laos1/531.9%
MaravarsTamil Ramanathapuram1/801.3%Dry Land Farmers
GarosGaro Tangail1/1200.8%Likely P77+

With K-M9+, unconfirmed but probable T-M70+: 56.6% of Kunabhis in Uttar Kannada, 32.5% of Kammas in Andhra Pradesh, 26.8% of Brahmins in Visakhapatnam, 25% of Kattunaiken in South India, 22.4% of Telugus in Andhra Pradesh, 20% of Ansari in South Asia, of Poroja in Andhra Pradesh, 9.8% of Kashmiri Pandits in Kashmir, 8.2% of Gujars in Kashmir, 7.7% of Siddis in Andhra Pradesh, 5.5% of Adi in Northeast India, 5.5% of Pardhans in Adilabad, 5.3% of Brahmins in Bihar, 4.3% of Bagata in Andhra Pradesh, 4.2% of Valmiki in Andhra Pradesh, of Brahmins in Maharashtra, 3.1% of Brahmins in Gujarat, 2.9% of Rajput in Uttar Pradesh, 2.3% of Brahmins in Peruru, and 1.7% of Manghi in Maharashtra.
Also in Desasth-Brahmins in Maharashtra and Chitpavan-Brahmins in Konkan, Chitpavan-Brahmins in Konkan.

Central Asia & East Asia

Unconfirmed but probable T-M70+: 2% of Hui in Liaoning, and 0.9% of Bidayuh in Sarawak.

Ancient DNA

Abel Beth Maacah

Abel Beth Maacah 2201 was a man with Y-DNA T-CTS2860 who lived between 1014 - 836 BCE during the Levant Iron Age and was found in the region now known as Abel Beth Maacah, Metula,Israel. At the Iron Age layer which also produces a Yahwistic inscription on a pottery jar from the biblical site of Abel-beth-maachah, which bears a faint Hebrew inscription of the name "Benayau".

Ancient Egypt

Egyptian mummy 2516 was a man who lived between 798 - 591 BCE during the Third Intermediate Age and was found in the region now known as Egypt. He is wearing a curly wig, a shabti made of multicoloured wood and a multicoloured wesekh-collar. There is an inscription, encircling the entire body in horizontal lines, with the text of Chapter VI of the Book of the Dead. The Ancient Egyptian was under T-Y6671, the saharan offshoot of T-L208 ultimately derived from T-M70.

Ancient Nubia

Multiple Nubians from Kulubnarti site were found to be of the Haplogroup T lineage, same as the ancient Egyptian clade. The Kulubnarti Nubians had ~43% Nilotic-related ancestry with the remaining ancestry consistent with being introduced through Egypt and ultimately deriving from an ancestry pool like that found in the Bronze and Iron Age Levant. It is hypothesized "T" lineage originated or evolved in the Levant, and became Saharan Pastoralists via their spread into Africa during the Neolithic. T-Y6671 is associated with this spread. This falls in line perfectly when considering the Levantine-like DNA that the Nubians harbor in concomitance to T-Y6671. The Nubian samples include I6328, I6340 & I19140. These Nubians lived during 700 - 990 CE and were found in "R and S Cemeteries", where E & J haplogroup was buried amongst these "T" individuals.

Neolithic North Africans

During the Neolithic Era a new ancestry from the Levant appears in the Maghreb, coinciding with the arrival of pastoralism in the region, and all three ancestries blend together during the Late Neolithic. This places Haplogroup T as a pastoralist lineage, and due to its circumstances, is associated with Levantine expansion,spreading Afro-Asiatic languages, eventually morphing into Saharan Pastoralists and spreading Afro-Asiatic languages. Sample SKH003 and SKH002 were Neolithic local Northwest African and differentiated from older Northwest Africans exactly due to an influx of Levantine PPNB ancestry. This ancestry was introduced with this new Y-chromosome haplogroup, and is very clearly a Male dominated migration, as only Y-chromosome lineages were replaced, and no mtdna was introduced. Unlike the earlier expansion of Anatolian Neolithic / Early European farmer dna, which were maternally lead migrations.

Peki'in Cave, Israel

A 2018 study conducted by scholars from Tel-Aviv University, the Israel Antiquities Authority and Harvard University had discovered that 22 out of the 600 people who were buried in Peki'in cave from the Chalcolithic Period were of both local Levantine and Zagros area ancestries, or as phrased in the paper itself: "Ancient DNA from Chalcolithic Israel reveals the role of population mixture in cultural transformation," the scientists concluded that the homogeneous community found in the cave could source ~57% of its ancestry from groups related to those of the local Levant Neolithic, ~26% from groups related to those of the Anatolian Neolithic, and ~17% from groups related to those of the Iran Chalcolithic.". The scholars noted that the Zagros genetic material held "Certain characteristics, such as genetic mutations contributing to blue eye color, were not seen in the DNA test results of earlier Levantine human remains MTDNA blue-eyed, fair-skinned community didn't continue, but at least now researchers have an idea why. "These findings suggest that the rise and fall of the Chalcolithic culture are probably due to demographic changes in the region".
We find that the individuals buried in Peqi'in Cave represent a
relatively genetically homogenous population. This homogeneity
is evident not only in the genome-wide analyses but also in the
fact that most of the male individuals belong to
the Y-chromosome Haplogroup T (Y-DNA), a
lineage thought to have diversified in the Near East. This
finding contrasts with both earlier
Levantine populations, which were dominated by Haplogroup E (Y-DNA),
and later Bronze Age individuals, all of whom belonged to Haplogroup J (Y-DNA).

Ancient city of Ebla

In the ancient city of Ebla in Syria in the Bronze Age, one individual was found belonging to haplogroup T-L162.

Alalakh Amorite city-state

One individual from Alalakh who lived circa 2014-1781 BC, belonged to haplogroup T-CTS11451.

Notable haplogroup members

Elite endurance runners

Possible patterns between Y-chromosome and elite endurance runners were studied in an attempt to find a genetic explanation to the Ethiopian endurance running success. Given the superiority of East African athletes in international distance running over the past four decades, it has been speculated that they are genetically advantaged. Elite marathon runners from Ethiopia were analysed for K* which according to the previously published Ethiopian studies is attributable to the haplogroup T.
According to further studies, T1a1a* was found to be proportionately more frequent in the elite marathon runners sample than in the control samples than any other haplogroup, therefore this y-chromosome could play a significant role in determining Ethiopian endurance running success. Haplogroup T1a1a* was found in 14% of the elite marathon runners sample of whom 43% of this sample are from Arsi province. In addition, haplogroup T1a1a* was found in only 4% of the Ethiopian control sample and only 1% of the Arsi province control sample. T1a1a* is positively associated with aspects of endurance running, whereas E1b1b1 is negatively associated.

House of Khalifa

The ruling family of the Kingdom of Bahrain is the House of Khalifa is confirmed West Asian Y-DNA Haplogroup T-L206 subclade of P77*.
The house belongs to the Utab tribe, which is part of the larger Anizah tribal confederation, that migrated from Central Arabia to Kuwait and then ruled all of Qatar. In 1999, Hamad bin Isa Al Khalifa became the Emir of Bahrain and proclaimed himself the King of Bahrain in 2002.
The T-FT364053 haplogroup of the house was determined by DNA testing of descendants in the T-Arab Y DNA Haplogroup Project on Family Tree DNA and other Arab world projects.

Thomas Jefferson

A notable member of the T-M184 haplogroup is American President Thomas Jefferson. The Y-chromosomal complement of the Jefferson male line was studied in 1998 in an attempt to resolve the controversy over whether he had fathered the mixed-race children of his slave Sally Hemings. A 1998 DNA study of the Y chromosome in the Jefferson male line found that it matched that of a descendant of Eston Hemings, Sally Hemings' youngest son. This confirmed the body of historical evidence, and most historians believe that Jefferson had a long-term intimate liaison with Hemings for 38 years, and fathered her six children of record, four of whom lived to adulthood. In addition, the testing conclusively disproved any connection between the Hemings descendant and the Carr male line. Jefferson grandchildren had asserted in the 19th century that a Carr nephew had been the father of Hemings' children, and this had been the basis of historians' denial for 180 years.
Jefferson's paternal family traced back Wales, where T is incredibly rare, as it is less than <1% throughout Britain. A couple of British males with the Jefferson surname have been found with the third president's type of T, reinforcing the likelihood that his immediate paternal ancestry was British.
Family Tree DNA, found that the Jefferson T patrilineage belongs to T-BY78550 a subclade of T-PF7444 which is likely of MENA Middle Eastern North African Origins. Spencer Wells who led The Genographic Project places his origin to Canaan

Nomenclatural history

Prior to 2002, there were in academic literature at least seven naming systems for the Y-Chromosome Phylogenetic tree. This led to considerable confusion. In 2002, the major research groups came together and formed the Y-Chromosome Consortium. They published a joint paper that created a single new tree that all agreed to use. Later, a group of citizen scientists with an interest in population genetics and genetic genealogy formed a working group to create an amateur tree aiming at being above all timely. The table below brings together all of these works at the point of the landmark 2002 YCC Tree. This allows a researcher reviewing older published literature to quickly move between nomenclatures.
YCC 2002/2008 YCC 2002 YCC 2005 YCC 2008 YCC 2010r ISOGG 2006ISOGG 2007ISOGG 2008ISOGG 2009ISOGG 2010ISOGG 2011ISOGG 2012ISOGG 2013
T-M18426VIII1U25Eu16H5FK*KTTK2K2TTTTTT
K-M70/T-M7026VIII1U25Eu15H5FK2K2TT1K2K2TTTT1T1aT1a
T-P7726VIII1U25Eu15H5FK2K2T2T1a2K2K2T2T2T2a1T1a1bT1a1a1T1a1a1

Original research publications

The following research teams per their publications were represented in the creation of the YCC Tree.
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