Haplogroup O-M175


Haplogroup O, also known as O-M175, is a human Y-chromosome DNA haplogroup. It is primarily found among populations in Southeast Asia and East Asia. It also is found in various percentages of populations of the Russian Far East, Siberia, Central Asia, Caucasus, Crimea, Ukraine, Iran, South Asia, Oceania, Madagascar and the Comoros. Haplogroup O is a primary descendant of haplogroup NO-M214.
The O-M175 haplogroup is very common amongst males from East and Southeast Asia. It has two primary branches: O1 and O2 . O1 is found at high frequencies amongst males native to Southeast Asia, Taiwan, the Japanese Archipelago, the Korean Peninsula, Madagascar and some populations in southern China and Austroasiatic speakers of India. O2 is found at high levels amongst Han Chinese, Tibeto-Burman populations, Manchu, Mongols, Koreans, Vietnamese, Filipinos, Japanese, Thais, Polynesians, Miao people, Hmong, the Naiman tribe of Kazakhs in Kazakhstan, Kazakhs in the southeast of Altai Republic, and Kazakhs in the [Ili Kazakhs in China|Kazakh Autonomous Prefecture|Ili area] of Xinjiang.

Origins

Haplogroup O-M175 is a descendant haplogroup of Haplogroup NO-M214, and first appeared according to different theories either in Southeast Asia or East Asia approximately 40,000 years ago.
Haplogroup O-M175 is one of NO-M214's two branches. The other is Haplogroup N, which is common throughout North Eurasia.

Distribution

This haplogroup appears in high to moderate frequencies in most populations in both East Asia and Southeast Asia, and it is almost exclusive to that region. It is almost nonexistent in Western Siberia, Western Asia, Europe, most of Africa, India and the Americas, where its presence may be the result of recent migrations. However, certain O subclades do achieve significant frequencies among some populations of Central Asia, South Asia, and Oceania. For example, one study found it at a rate of 67% among the Naimans, a tribe in Kazakhstan, even though the rate among Kazakhs in general is only about 3.3% to 10.8%. Karafet et al. have assigned the Y-DNA of 46.2% of a sample of Papuan from Pantar Island to haplogroup NO-M214; considering their location in the Southeast Archipelago, all or most of these individuals should belong to haplogroup O-M175.
An association with the spread of Austronesian languages in late antiquity is suggested by significant levels of O-M175 among island populations of the South Pacific and Indian Ocean, including the East African littoral. For example, Haplogroup O-M50 has even been found in Bantu-speaking populations of the Comoros along 6% of O-MSY2.2(xM50), while both O-M50 and O-M95(xM88) occur commonly among the Malagasy people of Madagascar with a combined frequency of 34%. O-M175 has been found in 28.1% of Solomon Islanders from Melanesia. 12% of Uyghurs, 6.8% of Kalmyks, 6.2% of Altaians, 4.1% of Uzbeks on average but Uzbeks from Bukhara 12.1%, Karakalpaks 11.4%, Sinte 6.7% and 4.0% of Buryats.
In the Caucasus region it has been found in the Nogais 6% but 5.3% in the Karan Nogais, it is also found in the Dargins of Dargwa speakers at 2.9%. In the Iranic population, it is found in Iranian at 6.3%, 8.9% of Tajiks in Afghanistan 4.2% in the Pathans in Pakistan but 1% in Afghanistan, 3.1% in Burusho.
Haplogroup O-M175 ranges in various moderate to high frequencies in the ethnic minorities of South Africa. The frequency of this haplogroup is 6.14% in the Cape colored population, 18% in Cape Coloured Muslim, 38% in Cape Indian Muslims and 10% in Cape Other Muslims. It's found 11.5% in the Réunion Creole.
Haplogroup O-M175 had also been found in Latin America and Caribbean as a result of massive Chinese male migration from the 19th century. It was found in the Jamaicans at 3.8%, and in Cubans, 1.5%.
Haplogroup O-M175 has been found in 88.7% of Asian American. 1.6% in Hispanic American, White Americans 0.5%, and 0.3% in African American. Another study gives 0.5% African American.
Among the sub-branches of haplogroup O-M175 are [Haplogroup Haplogroup O-M119|O-M119 (Y-DNA)|O-M119], O-M268, and [Haplogroup Haplogroup O-M122|O-M122 (Y-DNA)|O-M122].
Y Haplogroup O3-M122 makes up the majority of Jadoon's males, the same haplogroup carried by the majority of Han Chinese. 82.5% of Jadoon men carrying Q-MEH2 and O3-M122 which are both of East Asian origin. O3-M122 was absent in the Sayyid population and appeared in low numbers among Tanolis, Gujars and Yousafzais. There appears to be founder affect in the O3-M122 among the Jadoon. 76.32% of Jadoon men carry O3-M122 while 0.75% of Tanolis, 0.81% of Gujars and 2.82% of Yousafzais carry O3-M122.
Russians in China East Asian haplogroup O made up 58% of their Y haplogroup. O3-M122 specifically made up 47% of the Russian sample. The East Asian Y haplogroup O3-M122 was found in 47% of Russian males in China. In another test the East Asian paternal Y Haplogroup O made up 58% of Russian males samples in China.
Haplogroup O was found in 1%-1.2% of Persians in one sample.
O3-M122 is the commonly shared genetic signature of Sino-Tibetan speaking ethnicities.

O-M175*

A broad survey of Y-chromosome variation among populations of central Eurasia found haplogroup O-M175 in 31% of a sample of Koreans and in smaller percentages of Crimean Tatars, Tajiks, Uyghurs, Uzbeks, and Kazakhs . However, nearly all of the purported Korean O-M175 Y-chromosomes may belong to Haplogroup O-M176, and later studies do not support the finding of O-M175* among similar population samples. The reported examples of O-M175 Y-chromosomes that have been found among these populations might therefore belong to Haplogroup O-M268*(xM95,M176) or Haplogroup O-M176.
A study published in 2013 found O-M175 Y-DNA in 5.5% Iranians from Teheran, 5.4% Tajiks from Badakhshan Province of Afghanistan, and 1/97 Mongols from northwest Mongolia, while finding O-M176 only in 1/20 Mongols from northeast Mongolia.

O-F265 (O1)

O1a-M119 and O1b-M268 share a common ancestor, O1-F265 approximately 33,181 YBP. O1-F265, in turn, coalesces to a common ancestor with O2-M122 approximately 33,943 YBP. Thus, O1-F265 should have existed as a single haplogroup parallel to O2-M122 for a duration of approximately 762 years before breaking up into its two extant descendant haplogroups, O1-MSY2.2 and O1b-M268.
O-M119 (O1a)
O-M119 is found frequently in Austronesian-speaking people, with a moderate distribution in southern and eastern Chinese and Kra-dai peoples.
O-M268 (O1b)
O-K18 Naxi
  • * O-CTS4040
  • ** O-MF56251 Observed sporadically in China, Thailand, Vietnam
  • ** O-Page59/CTS10887 Found among North Han Chinese, East Han Chinese, South Han Chinese
  • *** O-F4070
  • **** O-MF106398 Observed sporadically in China
  • **** O-F779/F993/F3135 China, Vietnam, Qatar
  • ***** O-MF107014 Observed sporadically in China
  • ***** O-CTS5160/MF61620 China
  • ***** O-F2064/F1759 China, Singapore, Vietnam, Korea
  • ****** O-PH2797/CTS1127 China
  • ******* O-MF66989 Observed sporadically in China
  • ******* O-PH2506 China, Japan
  • ****** O-Y148532 China, Afghanistan
  • ****** O-Y239146/MF31164 Singapore, Taiwan
  • ****** O-Y47392/MF17288 China
  • ****** O-BY182144/Y157814 China, Taiwan
  • ******* O-MF16472/MF148514 China
  • ******** O-MF184575 China
  • ******* O-ACT3266/Y149401 China
  • ****** O-PH4822 China
  • *** O-F417/M1654/CTS469 Japan
  • **** O-F840/F1247/F1378
  • ***** O-Y185784 Singapore
  • ***** O-CTS250
  • ****** O-F2760 China
  • ****** O-FGC30081 China
  • **** O-CTS1451 Korea, China
  • *** O-CTS9996/PF4341 Philippines
  • **** O-PF4341* Hunan
  • **** O-CTS298 China
  • ***** O-F16308 China
  • ***** O-MF2931 China
  • **** O-MF14344 China
  • ***** O-F1356 China, Vietnam, South Korea
  • * O-PK4
  • **O-F838 Found in about 1.4% of Han Chinese
  • *** O-CTS3857
  • **** O-F714
  • ***** O-F2081 Shaanxi
  • ***** O-F3357
  • ****** O-F977 Liaoning
  • ******* O-F1199 Beijing, Sichuan
  • ******** O-ACT7133 Chongqing Liwutu, Jiangxi
  • ******** O-Y168698 Sichuan, Anhui
  • ****** O-CTS4052/CTS5664 Hunan
  • ******* O-Y155291 Anhui, medieval Turk from the oasis of Otrar
  • ******* O-CTS2452 Guangxi, Hunan
  • **** O-F15640/F11242 Fujian, Singapore, Spain
  • ***** O-F16212
  • ****** O-F21414 Fujian
  • ******* O-F15452
  • *** O-F15300 Chongqing, Guizhou, Anhui
  • **** O-F14826 Hunan
  • ***** O-MF45275 Guangxi
  • ***** O-Y34152
  • ****** O-MF43808 Sichuan, Hunan
  • ****** O-MF37500 Hunan
  • ******* O-MF43192 Sichuan, Hunan
  • ** O-M95
  • *** O-CTS350 China
  • **** O-CTS350* Japan
  • **** O-CTS10007/CTS926 China
  • *** O-M1310
  • **** O-Y172653/Y172877 Found in China and Japan
  • **** O-F1803/M1348 China
  • ***** O-ACT721/ACT1038 Found sporadically in China
  • ***** O-F789/M1283 Found in China, Vietnam, Cambodia, Singapore, Java, Borneo, Thailand, Laos, Myanmar, Bhutan, Bangladesh, India
  • ****** O-M1283* Lao Isan
  • ****** O-MF600645 Gansu, Sichuan, Hunan
  • ****** O-M1368 Singapore
  • ******* O-M1361
  • ******** O-MF611153 Found sporadically in China
  • ******** O-A22938 Vietnam, China
  • ********* O-F2194 China
  • ********* O-BY189914 China
  • ********* O-F16163 China, Taiwan
  • ********** O-F20644 China
  • *********** O-SK1646 China
  • ******** O-Y9322 China
  • ********* O-Y9325
  • ********** O-Z39485 China
  • ********** O-Z39490
  • *********** O-Z39490* Indonesia, Singapore
  • *********** O-B421 Sichuan, Singapore, Java, Madhya Pradesh, Saudi Arabia
  • ********** O-Y9033/B426 Laos, Thailand, Vietnam
  • *********** O-Y9033* Indonesia, India, Bangladesh
  • *********** O-M6661 Kinh in Ho Chi Minh City
  • *********** O-YP3930 Cambodia, Thailand, Laos
  • ***** O-F1252
  • ******O-SK1630/F5504 China
  • *******O-ACT5802
  • ********O-MF92614
  • ********O-F16061
  • *********O-MF286118 Found in two Han Chinese from Guangdong
  • *********O-F19607
  • **********O-SK1636 China
  • **********O-MF206811 China
  • **********O-MF91645 China
  • **********O-Y68888 Russia
  • **********O-F5506 Hmong
  • ****** O-F2924
  • ******* O-CTS5854
  • ******** O-Z23810
  • ********* O-CTS7399
  • ********** O-Y85641 Japan, China
  • ********** O-FGC19713/Y14026 Laos, Thailand, Vietnam
  • *********** O-FGC19707
  • ************ O-MF14427 China
  • ************ O-FGC19716 China
  • ************* O-FGC19718 China, Philippines
  • ************ O-Z23849 China
  • ********* O-CTS651/CTS10484 Thailand, Laos, Vietnam
  • ********** O-CTS10484* China
  • ********** O-PH4310 China, Singapore
  • ********** O-Z1018 China
  • ******** O-Z23781 China
  • ********* O-F4229 Singapore
  • ********** O-F809 China
  • *********** O-Z23795 China
  • ************ O-MF223122 China
  • ************ O-A22937
  • ************* O-BY50703
  • ************** O-MF169272 China
  • ************** O-MF226417 China
  • ************* O-MF61630 China
  • ************ O-F2517 China
  • ************* O-MF5664
  • ************* O-Z23790
  • ******* O-M111/M88 Found frequently among Vietnamese, Tai peoples, Lachi, Lô Lô, Hani-Akha, Bunu, She people, Cambodians, Kuy, Bru, and Htin, with a moderate distribution among Qiang, Bai, Yi, Bamar, Jingpo, Lahu, Tujia, Han Chinese, Miao, Pathen, Yao, Hlai, Taiwanese aborigines, the Philippines, Malaysia, Kalimantan, Java, Chamic-speaking peoples, and Kiribati
  • ******** O-M111/M88* Northern Thailand, Cambodia, Yunnan
  • ******** O-F2524
  • ********* O-F2524* Jiangsu
  • ********* O-F2346
  • ********** O-F2890 Thailand, Vietnam
  • *********** O-F2890* Ho Chi Minh City
  • *********** O-Z24048
  • ************ O-F18990 Jiangsu, Jiangxi, Xishuangbanna
  • ************ O-Z24050
  • ************* O-Z24050* Xishuangbanna
  • ************* O-ACT7172 Guangdong
  • ************* O-Y151536
  • ************** O-Z22028 Hunan, Anhui, Guangdong
  • ************** O-Y151537
  • *************** O-Y151537* Hunan
  • *************** O-ACT523
  • **************** O-ACT505 Guangdong
  • **************** O-ACT502 Jiangxi, Hunan, Guangxi
  • ********** O-F2758 Vietnam, Cambodia, Thailand, Laos, Yunnan
  • *********** O-F2758* China
  • *********** O-Z24083
  • ************ O-Z24083* Ho Chi Minh City
  • ************ O-Z24089
  • ************* O-SK1627/Z24091 Vietnam, Thailand, Laos
  • ************** O-Z24091* China, Vietnam
  • ************** O-Y26364
  • *************** O-Y26364* Thailand
  • *************** O-Y26370 China, Vietnam
  • ************* O-F923
  • ************** O-F923* Xishuangbanna, Ho Chi Minh City
  • ************** O-Z24154 Ho Chi Minh City
  • ************** O-CTS2022
  • *************** O-CTS2022* Ho Chi Minh City
  • *************** O-F3053 Thailand
  • **************** O-F3053* Cambodian
  • **************** O-F4383/F1399 Vietnam
  • ***************** O-F4383/F1399* China, Singapore
  • ***************** O-F14994 Vietnam, Singapore
  • ***************** O-F2415 Thailand, China O-M176
  • * O-K4: Found frequently among Koreans and with a moderate distribution among Japanese, Ryukyuans, Daurs, Evenks, Hezhe, Manchus, and Sibe. Also found sporadically among Han Chinese, Hui, Micronesians, Mongols, Thais, Uyghurs, Vietnamese, etc.
  • * O-47z: Found frequently among Japanese and Ryukyuans and with a moderate distribution among Koreans. Found sporadically among Manchus, Mongols, Han Chinese, Hui, Tujia, Vietnamese, etc.

O-M122 (O2)

Found frequently among populations of East Asia, Southeast Asia, and culturally Austronesian regions of Oceania, with a moderate distribution in Central Asia.
  • O-M122
  • *O-CTS1754 East & Southeast Asia
  • **O-F953/F1886 Philippines
  • ***O-F953* Ho Chi Minh City
  • ***O-F1024
  • ****O-F1024* Guangxi
  • ****O-Y29790 Liaoning, Fujian
  • *O-M324
  • **O-L465
  • ***O-CTS727
  • ****O-F915
  • *****O-F915* Shanxi
  • *****O-Y14462
  • ******O-Y14462* Guangdong
  • ******O-CTS1936 Beijing, Ho Chi Minh City
  • ******O-A18267
  • *******O-A18029 China
  • *******O-ACT6559 Hebei
  • ****O-CTS3709
  • *****O-CTS11209
  • ******O-CTS11209* Han from Hunan
  • ******O-L599 Han from Hunan, Kinh from Ho Chi Minh City, Tujia
  • *****O-F2640
  • ******O-F2640* Beijing
  • ******O-CTS11210
  • *******O-CTS11210* Han from Hunan
  • *******O-CTS2432 Han from Hunan
  • ***O-JST002611/CTS2483
  • ****O-CTS2483* China, Japan, Philippines
  • ****O-CTS10573 Beijing, Sichuan, Henan, Jiangsu
  • *****O-CTS10573* Beijing, Dai from Xishuangbanna
  • *****O-MF7367
  • ******O-MF7367* Dai
  • ******O-MF9367 South Korea, Japan
  • ****O-F18
  • *****O-CTS498 China, Japan
  • *****O-F449 Azerbaijan
  • ******O-F1266, F2016, F4267 Beijing, Jilin, Shanxi, Jiangsu, Shanghai, Guangdong
  • ******O-F238
  • *******O-FGC62457
  • ********O-CTS679 Hubei, Fujian
  • ********O-MF19356 Guizhou, Chongqing, Jiangxi
  • *******O-F134
  • ********O-F134*
  • ********O-F1894 Beijing, Shanghai, Liaoning, Jilin, Hebei, Shanxi, Zhejiang, Guangdong
  • *********O-F1894* Hunan, Beijing
  • *********O-MF18144
  • **********O-MF18144* Shandong
  • **********O-F18567 South Korea, Japanese
  • ********O-MF8794
  • *********O-MF8794* China
  • *********O-MF1155
  • **********O-MF1155* Anhui, Myanmar
  • **********O-MF1157 Guangdong, Jiangxi, Russia
  • *****O-F117
  • ******O-F117* Fujian
  • ******O-F11
  • *******O-F11* Gansu, Japanese
  • *******O-F930 Beijing, Armenia, Inner Mongolia, Hebei, Shaanxi, Shandong, Zhejiang, Hubei
  • ********O-F2685 Beijing, Shanghai, Fujian, Guangdong
  • *******O-BY169374
  • ********O-BY169374* Zhejiang
  • ********O-SK1692
  • *********O-SK1692* Guangdong
  • *********O-MF55116 Pakistan
  • *******O-F539 Beijing, Shanghai, Jiangsu, Zhejiang, Jiangxi, Guangdong, Yunnan
  • ********O-F539*
  • ********O-F319 Vietnam, Hunan
  • *******O-CTS12877
  • ********O-F16635
  • *********O-F16635* South Korea
  • *********O-ACT3882 Hubei, Hunan
  • ********O-CTS257 Zhejiang, Shanghai, Guangdong, Beijing, Fujian, Heilongjiang, Hunan
  • *********O-CTS257* South Korea, Japan
  • *********O-F16834
  • **********O-F16834* Vietnam
  • **********O-Y166692 Lahu, Yi
  • *******O-Y29837
  • ********O-Y29837* Hebei, Fujian
  • ********O-MF8526
  • *********O-MF8526* Sichuan, Zhejiang
  • *********O-Y135777 China
  • *******O-BY36917 Japan
  • ********O-F270 Beijing, Hebei, Shandong
  • *********O-F270* Hezhen
  • *********O-F1272 Jiangsu
  • *******O-F4062 Beijing, Shanghai, Guangdong, Jiangsu, Shandong, Shaanxi, Chongqing, Heilongjiang, Liaoning, Henan, Hubei, Hunan, Zhejiang
  • ********O-Y15976 China, Japan, Korea, Pakistan, Vietnam
  • *********O-Y15976* Zhejiang, Vietnam
  • *********O-Y26383
  • **********O-Y26383* Hubei
  • **********O-SK1686
  • ***********O-SK1686* Pakistan
  • ***********O-Y46851 South Korea
  • ********O-FGC54474
  • *********O-FGC54474* Skeleton exhumed from a mass grave located in Nomgon, Ömnögovi, Mongolia
  • *********O-FGC54505 China, Japan
  • *********O-MF8460 Anhui
  • *********O-MF9523 South Korea
  • *******O-F971 Beijing, Shanghai, Hubei, Guangdong
  • ********O-F971* Sichuan, Zhejiang, Jiangxi, Japan
  • ********O-Y23813
  • *********O-Y23813* Guangdong
  • *********O-Y23809
  • **********O-Y23809* Guangdong
  • **********O-BY170727
  • **********O-Y23907 Guangdong
  • *******O-F632
  • ********O-F632* Beijing
  • ********O-F16340 Zhejiang
  • ********O-F133 China, Bulgaria
  • *********O-F133* Beijing
  • *********O-Y20951
  • **********O-Y20951* Anhui
  • **********O-Y20932 Anhui, Beijing
  • *********O-F17
  • **********O-F377 China, France
  • ***********O-CTS7789 Han Chinese
  • ************O-CTS7789* Beijing, Guangdong
  • ************O-F1095
  • *************O-F1095* Fujian
  • *************O-Z25097
  • **************O-Z25097* Hunan
  • **************O-Z7776 Hunan, Guangdong
  • ***********O-CTS7501
  • ************O-CTS7501* Zhejiang, Hunan
  • ************O-CTS1621
  • *************O-CTS1621* Hubei, Hunan
  • *************O-Y147084 Jiangxi
  • **********O-F793 Beijing, Tianjin, Shandong, Shanxi, Shaanxi, Xinjiang, Jiangsu, Shanghai, Hubei, Sichuan, Guangdong
  • ***********O-F793* Sichuan, Guangdong, Guizhou, Hunan, Jiangsu, French Polynesia, She, North China
  • ***********O-Y150436
  • ************O-Y150436* Jiangxi
  • ************O-Y150331 Jiangxi, Yunnan
  • ***********O-Y141213 Jilin, Liaoning, Henan, Shandong, Jiangsu
  • **O-P201
  • ***O-M188
  • ****O-M188* Korea
  • ****O-CTS800
  • *****O-CTS800* Beijing, Japan
  • *****O-CTS1602 Han from Fujian
  • ****O-CTS445
  • *****O-CTS201 Korea
  • ******O-M159 China, Taiwan, Cambodia, Malaysia, Singapore
  • *******O-FTA21663/O-MF22947 China, Saudi Arabia
  • *******O-CTS3994
  • ********O-MF125236 China
  • ********O-Z25482
  • *********O-Z25518 China
  • **********O-Z35182
  • ***********O-Z35182* China
  • ***********O-MF2541 Fujian, Taiwan, Cambodia
  • **********O-CTS5032
  • ***********O-CTS5032* Hunan
  • ***********O-CTS10690 Hunan
  • ******O-MF18110/FGC50590 China
  • *******O-MF109844
  • ********O-MF106775 China
  • ********O-MF106730 China
  • *******O-FGC50661 China
  • ********O-MF56709
  • *********O-MF213370 Found sporadically in Yangzhou, Shanghai, Liangshan
  • *********O-MF56898 China
  • ********O-FGC50643/MF15475 China
  • *********O-MF56474 China
  • *********O-FGC50649
  • **********O-Y169670/O-MF14256 China
  • ***********O-MF50824
  • ************O-MF106512 China
  • ************O-MF88959/O-Y150315 South Korea, China
  • ***********O-MF14135/O-Z12303 China
  • ************O-MF238642
  • *************O-MF37094 China
  • *************O-Y169696/O-MF15693 China
  • *************O-MF18577/O-MF18626 China, Kazakhstan, Thailand
  • **********O-FGC50558 Japan, Korea
  • ***********O-FGC50558* Korea, Japanese
  • ***********O-Y165513 Korea
  • ***********O-FGC50535 Korea
  • *****O-M7 Found frequently among human remains associated with the Neolithic Daxi culture and modern Hmong–Mien, Katuic, and Bahnaric peoples, with a moderate distribution among Han Chinese, Buyei, Bai, Mosuo, Tibetans, Qiang, Oroqen, Tujia, Thai, Orang Asli, western Indonesians, Malaysians, Vietnamese, and Atayal.
  • ******O-MF106687 China
  • ******O-Z25245
  • *******O-MF9858/O-Z6157 China, Thailand
  • ********O-MF9896 China
  • ********O-PH204 China, Taiwan
  • *******O-Y26422
  • ********O-Y26395/O-SK1713 Taiwan
  • ********O-F21738 Philippines, Malaysia, Indonesia, Taiwan
  • *******O-F1276
  • ********O-F1863
  • *********O-MF107102 China
  • *********O-MF56735 China
  • *********O-MF36531 China
  • **********O-Y13816
  • ***********O-MF109664 China
  • ***********O-MF36502 Guangdong
  • *********O-F1134
  • **********O-MF35799/O-Y94171 Thailand, China
  • **********O-F1262/O-Y173492 China
  • *********O-FT303223/O-MF106843/O-F15314/O-F20756 China, Thailand
  • *********O-Z25288/O-Z25293 Vietnam
  • ********O-CTS6489
  • *********O-MF106428/O-Y94472/O-FTB23660 Thailand, Vietnam, China
  • *********O-F1275 Guangxi
  • **********O-MF15199/O-FTA25885
  • ***********O-F20472
  • ************O-FTB23785 Thailand, Vietnam
  • ************O-F17410/O-F18833/O-MF122643/O-BY177553 Thailand
  • *************O-MF106415/O-MF111486/O-BY122399 Thailand, China
  • *************O-Y127482/O-F15988 Thailand
  • **************O-F23936 Thailand, Singapore
  • ***************O-FTB24349/O-FTB25408 Thailand
  • **************O-F26466 Thailand
  • ***************O-F15259/O-Y238116 Singapore
  • ****************O-F19496
  • *****************O-F19315 Singapore
  • *****************O-F14817 Singapore, Thailand, Indonesia
  • ***********O-MF6534/O-MF58872/O-BY27925/O-Y23477 Thailand, Singapore, China
  • ***********O-CTS6579
  • ************O-CTS123/O-F22573/O-MF48275 China
  • ************O-F14832/O-F15788/O-Y208219 China, Thailand
  • **********O-Z25411
  • ***********O-ACT1126/O-Y140772/O-F1289 China, Thailand
  • ***********O-Z25398
  • ************O-F22005/O-Z25400 Thailand, Vietnam, China
  • ************O-F1100/O-Y37861 Hunan
  • *************O-MF17697 Laos, Thailand, China
  • *************O-F1234/O-Y37855
  • **************O-Y185160/O-MF36985 Hebei, Beijing, Sichuan, Shaanxi, Guangxi, Zhejiang, Shandong, Ningxia, Inner Mongolia, Hubei, Jiangxi
  • **************O-FGC71370
  • ***************O-MF193618 Sichuan, Zhejiang, Shandong, Anhui, Hunan, Hubei, Fujian, Philippines
  • ***************O-F14904/N5 Ningxia, Hmong, She, Iu Mien, Quebec. Huang et al. found that this is the most common Y-chromosome haplogroup among many Hmongic-speaking ethnic groups, with a frequency of 47.1% among the Guangxi Miao.
  • ***O-P164
  • ****O-F996/F3237
  • *****O-A16433 Heilongjiang
  • ******O-MF56976 Anhui
  • ******O-Y125645
  • *******O-Y149134 Anhui, Henan
  • *******O-PF5573
  • ********O-PF5573* Guangdong
  • ********O-Y101407 Korea
  • ********O-MF14338
  • *********O-MF14338* Tatarstan
  • *********O-Y153434
  • **********O-Y153434* Shandong
  • **********O-MF76414 Chongqing, Skeleton exhumed from a mass grave located in Nomgon, Ömnögovi, Mongolia
  • *****O-F871
  • ******O-F706 Philippines, China, Cambodia, Thailand
  • *******O-F1010 Thailand
  • ********O-F1010* Zhejiang
  • ********O-Y170907
  • *********O-Y170907* South Korea
  • *********O-Y171445 Zhejiang, Japan
  • *******O-AM01750/AM01861/B451 Singapore, Indonesia, Philippines
  • ********O-AM01861* Spain
  • ********O-B450 Philippines, Indonesia, Singapore
  • *********O-BY157019 Mexico, Micronesia
  • ********O-A16139 Philippines, USA
  • ******O-F2472
  • *******O-F4110 Thailand
  • ********O-F4110* Fujian
  • ********O-MF1510
  • *********O-MF1510* Tonga
  • *********O-ACT3918 Zhejiang
  • *******O-F4124
  • ********O-F15739
  • *********O-F14875 Vietnam, Xinjiang
  • ********O-F26088/JST008425p6 Japan, Hunan
  • *********O-A16609
  • **********O-MF1232 Henan, Fujian
  • ****O-M134: Found frequently among speakers of Sino-Tibetan languages, among members of the Kazakh Naiman tribe with a moderate distribution throughout East Asia and Southeast Asia.
  • *****O-Y20/PAGES00125 Poland
  • ******O-F1725
  • *******O-F1725* Ho Chi Minh City
  • *******O-Y31255
  • ********O-FGC61200
  • *********O-FGC61200* Henan
  • *********O-FGC61168 Sichuan
  • ********O-A16615/CTS11009
  • *********O-Y85894
  • **********O-Y85894* Fujian
  • **********O-Y19010 Shandong
  • *********O-CTS682/CTS2272
  • **********O-BY146675 Korean
  • **********O-BY161237 Japan
  • ******O-Y12/F314
  • *******O-Y12* Beijing
  • *******O-CTS2643/CTS11192
  • ********O-CTS53
  • *********O-CTS53* China
  • *********O-CTS6373 Hunan, Xishuangbanna
  • ********O-F876
  • *********O-F876* Henan, Shaanxi
  • *********O-Y29829 Liaoning
  • ********O-F275
  • *********O-F275* Henan, Shandong
  • *********O-F14411/CTS1011 Japan
  • **********O-Z2037 Beijing
  • **********O-CTS2815
  • ***********O-CTS2815* South Korea
  • ***********O-F700
  • ************O-F700* Hunan, Liaoning, Japan
  • ************O-F14475 Hunan
  • *************O-F2505
  • **************O-F2505* Hebei
  • **************O-MF194 Korean, Ho Chi Minh City
  • ********O-F634
  • *********O-F634* Shandong
  • *********O-F209 Beijing
  • **********O-F1391
  • ***********O-F1391* Zhejiang
  • ***********O-F4117 Jiangxi, Sichuan
  • **********O-A5034
  • ***********O-MF357
  • ************O-ACT1039
  • ************O-MF308 Korean
  • ***********O-CTS4266
  • ************O-F14839 Henan
  • ************O-Z26108
  • *************O-Y30135
  • **************O-Y30135* Beijing
  • **************O-F2173
  • ***************O-F2173* Sichuan
  • ***************O-F1458 Shandong, Anhui
  • *************O-Z26156
  • **************O-Z26156* Ho Chi Minh City
  • **************O-BY102696
  • ***************O-BY102696* Korean
  • ***************O-BY61500 Guangdong
  • ********O-CTS3776/F2887
  • *********O-CTS3776* Beijing, Fujian, Japan
  • *********O-CTS3763
  • **********O-CTS4601 Fujian
  • **********O-Y7110
  • ***********O-Y13527
  • ************O-Y13527* Beijing
  • ************O-FGC16888/Z38720 Taiwan
  • *************O-MF805 Guizhou, South Korea
  • *************O-Y22112
  • **************O-FGC16885/FGC16887 Kazakhstan
  • ***********O-Y7111
  • ************O-MF13415 China
  • ************O-F4249/SK1769 China, South Korea
  • *************O-F4249* Jiangsu
  • *************O-FGC23868
  • **************O-FGC23868* Mongol
  • **************O-FGC23859 Jiangsu
  • ************O-CTS335 Zhejiang, Jiangsu, Gansu
  • *************O-CTS335* Japan, Xibo
  • *************O-Y142071
  • **************O-Y142071* Anhui
  • **************O-CTS3856
  • ***************O-CTS3856* Beijing
  • ***************O-CTS5741
  • ****************O-CTS1174 Hunan
  • ****************O-F15948 Jiangxi, Thailand
  • *****O-M117/PAGE23
  • ******O-MF1380/CTS4960 China, Korea, Japan, Indonesia
  • *******O-MF1388
  • ********O-MF1388* North China, Japan
  • ********O-Y139283 Anhui, Guangdong
  • *******O-MF2636
  • ********O-MF2636* Chongqing
  • ********O-CTS374 Jiangsu, Beijing
  • ******O-M133/M1706 Shandong
  • *******O-M1706* Japan
  • *******O-YP4864
  • ********O-YP4864* Korean
  • ********O-Z44076 Shandong, Philippines
  • *******O-CTS7634
  • ********O-CTS7634* Fujian, Beijing, Mongolia, Russia
  • ********O-F2188
  • *********O-MF23641
  • **********O-MF23641* China
  • **********O-Y169190 Fujian
  • *********O-Y29861 Henan, Fujian, Ho Chi Minh City
  • *********O-F3039 Shandong, Ho Chi Minh City, Philippines
  • *******O-M1726
  • ********O-A9457
  • *********O-A9457*
  • *********O-MF15246 Zhejiang, Korean
  • *********O-F17158
  • **********O-F15367
  • ***********O-F15367* Guangdong
  • ***********O-Y174543 Hunan, Japan
  • ********O-CTS9678
  • *********O-CTS9678* Tianjin
  • *********O-M1513
  • **********O-M1513* Shandong
  • **********O-F18746 Fujian
  • *********O-Z39663 Russia, Liaoning, Shandong, Fujian
  • *******O-A9459
  • ********O-F6800
  • *********O-F6800* Shanxi
  • *********O-Y8389
  • **********O-Y8389* Japan
  • **********O-Y9144
  • ***********O-Y9144* Guangdong
  • ***********O-Y8387
  • ************O-Y8387* Shanghai, Chongqing
  • ************O-CTS5481 Japan, Korean
  • ********O-F14249
  • *********O-F14249* Liaoning
  • *********O-ACT4506 Mongol, Guangdong
  • *********O-Y81597 South Korea, North Korea
  • *********O-F14347
  • **********O-F14347* Beijing
  • **********O-CTS5063
  • ***********O-CTS5063* Sichuan, Jiangxi, Fujian
  • ***********O-Z44927
  • ************O-Z44927* Anhui
  • ************O-MF21798 Zhejiang, Xishuangbanna
  • ***********O-CTS1154
  • ************O-CTS1154* Fujian, Guangxi
  • ************O-Z25902
  • *************O-Z25902* Hebei, Fujian
  • *************O-Z42620 Fujian, Ho Chi Minh City
  • ********O-F438 Japan
  • *********O-F438* Hunan
  • *********O-Y17728
  • **********O-Y17728* Beijing, Shandong
  • **********O-Y170157 Shandong, Korean
  • **********O-F1754
  • ***********O-F1754* Beijing
  • ***********O-F2137
  • ************O-F2137* Jiangsu
  • ************O-F1123 Jilin, Zhejiang
  • **********O-F316
  • ***********O-F316* Anhui
  • ***********O-CTS1304
  • ************O-CTS1304* Jiangsu, Shandong
  • ************O-F813
  • *************O-F813* Shandong, Jiangxi, Guangdong
  • *************O-Y138426 Shandong, Chongqing
  • *************O-Y154488 Hunan
  • *************O-Y20928 Henan
  • **************O-Y20928* Inner Mongolia, Beijing, Shandong, Anhui
  • **************O-Y174105
  • ***************O-Y174105* Henan
  • ***************O-Y137637 Shandong, Jiangsu
  • ********O-CTS1642
  • *********O-CTS1642* Bangladesh
  • *********O-CTS5308
  • **********O-CTS5308* Beijing
  • **********O-F20067
  • ***********O-F20067* Shannan
  • ***********O-M3873 Gansu, Shigatse
  • **********O-Y34065
  • ***********O-Y34065* Northeast India
  • ***********O-F16227 Shigatse, Xishuangbanna
  • **********O-MF1012
  • ***********O-MF1012* Nyingchi
  • ***********O-Y67478
  • ************O-Y67478*
  • ************O-CTS5672 Japanese
  • *********O-F14408
  • **********O-Y101264 Shigatse, Northeast India
  • **********O-F14422
  • ***********O-F14422* Northeast India, Myanmar
  • ***********O-A9463
  • ************O-A9463* Bangladesh
  • ************O-Y34067 Northeast India
  • ***********O-F14665 Xishuangbanna, Naxi
  • ***********O-Y7080
  • ************O-Y7080* Xishuangbanna, Guangdong
  • ************O-ACT1169 Guangdong, Fujian
  • ************O-Z26030
  • *************O-Z26030* Guangxi
  • *************O-F14479 ''Guangdong, Xishuangbanna, Ho Chi Minh City ''
    O-M324 (O2a)
O-F742 (O2b)

Language families and genes

Haplogroup O is associated with populations which speak Austric languages.
The following is a phylogenetic tree of language families and their corresponding SNP markers, or haplogroups, sourced mainly from and. This has been called the "Father Tongue Hypothesis" by George van Driem. It does not appear to account for O-M176, which is found among Japanese, Korean, and Manchurian males.

Phylogenetics

Phylogenetic history

Prior to 2002, there were in academic literature at least seven naming systems for the Y-Chromosome Phylogenetic tree. This led to considerable confusion. In 2002, the major research groups came together and formed the Y-Chromosome Consortium. They published a joint paper that created a single new tree that all agreed to use. Later, a group of citizen scientists with an interest in population genetics and genetic genealogy formed a working group to create an amateur tree aiming at being, above all, timely. The table below brings together all of these works at the point of the landmark 2002 YCC Tree. This allows a researcher reviewing older published literature to quickly move between nomenclatures.
YCC 2002/2008 ''''''YCC 2002 YCC 2005 YCC 2008 YCC 2010r ISOGG 2006ISOGG 2007ISOGG 2008ISOGG 2009ISOGG 2010ISOGG 2011ISOGG 2012
O-M17526VII1U28Eu16H9IO*OOOOOOOOOO
O-M11926VII1U32Eu16H9HO1*O1aO1aO1aO1aO1aO1aO1aO1aO1aO1a
O-M10126VII1U32Eu16H9HO1aO1a1O1a1aO1a1aO1a1O1a1O1a1aO1a1aO1a1aO1a1aO1a1a
O-M5026VII1U32Eu16H10HO1bO1a2O1a2O1a2O1a2O1a2O1a2O1a2O1a2O1a2O1a2
O-P3126VII1U33Eu16H5IO2*O2O2O2O2O2O2O2O2O2O2
O-M9526VII1U34Eu16H11GO2a*O2aO2aO2aO2aO2aO2aO2aO2aO2a1O2a1
O-M8826VII1U34Eu16H12GO2a1O2a1O2a1O2a1O2a1O2a1O2a1O2a1O2a1O2a1aO2a1a
O-SRY46520VII1U35Eu16H5IO2b*O2bO2bO2bO2bO2bO2bO2bO2bO2bO2b
O-47z5VII1U26Eu16H5IO2b1O2b1aO2b1O2b1O2b1aO2b1aO2b1O2b1O2b1O2b1O2b1
O-M12226VII1U29Eu16H6LO3*O3O3O3O3O3O3O3O3O3O3
O-M12126VII1U29Eu16H6LO3aO3aO3a1O3a1O3a1O3a1O3a1O3a1O3a1O3a1aO3a1a
O-M16426VII1U29Eu16H6LO3bO3bO3a2O3a2O3a2O3a2O3a2O3a2O3a2O3a1bO3a1b
O-M15913VII1U31Eu16H6LO3cO3cO3a3aO3a3aO3a3O3a3O3a3aO3a3aO3a3aO3a3aO3a3a
O-M726VII1U29Eu16H7LO3d*O3cO3a3bO3a3bO3a4O3a4O3a3bO3a3bO3a3bO3a2bO3a2b
O-M11326VII1U29Eu16H7LO3d1O3c1O3a3b1O3a3b1-O3a4aO3a3b1O3a3b1O3a3b1O3a2b1O3a2b1
O-M13426VII1U30Eu16H8LO3e*O3dO3a3cO3a3cO3a5O3a5O3a3cO3a3cO3a3cO3a2c1O3a2c1
O-M11726VII1U30Eu16H8LO3e1*O3d1O3a3c1O3a3c1O3a5aO3a5aO3a3c1O3a3c1O3a3c1O3a2c1aO3a2c1a
O-M16226VII1U30Eu16H8LO3e1aO3d1aO3a3c1aO3a3c1aO3a5a1O3a5a1O3a3c1aO3a3c1aO3a3c1aO3a2c1a1O3a2c1a1

Original Research Publications

The following research teams per their publications were represented in the creation of the YCC Tree.

Phylogenetic trees

ISOGG 2017 tree.
  • O
  • *O1
  • **O1a
  • ***O1a1
  • ****O1a1a
  • *****O1a1a1
  • ******O1a1a1a
  • *******O1a1a1a1
  • ********O1a1a1a1a
  • *********O1a1a1a1a1
  • **********O1a1a1a1a1a
  • ***********O1a1a1a1a1a1
  • ************O1a1a1a1a1a1a
  • *************O1a1a1a1a1a1a1
  • **************O1a1a1a1a1a1a1a
  • *************O1a1a1a1a1a1a2
  • *************O1a1a1a1a1a1a3
  • *************O1a1a1a1a1a1a4
  • *************O1a1a1a1a1a1a5
  • ************O1a1a1a1a1a1b
  • *************O1a1a1a1a1a1b1
  • ************O1a1a1a1a1a1c
  • *************O1a1a1a1a1a1c1
  • ************O1a1a1a1a1a1d
  • ************O1a1a1a1a1a1e
  • *************O1a1a1a1a1a1e1
  • ***********O1a1a1a1a1a2
  • *********O1a1a1a1a2
  • *******O1a1a1a2
  • ********O1a1a1a2a
  • *********O1a1a1a2a1
  • **********O1a1a1a2a1a
  • ********O1a1a1a2b
  • ******O1a1a1b
  • *******O1a1a1b1
  • *******O1a1a1b2
  • ********O1a1a1b2a
  • *********O1a1a1b2a1
  • *****O1a1a2
  • ******O1a1a2a
  • *******O1a1a2a1
  • ****O1a1b
  • ***O1a2
  • ****O1a2a
  • *****O1a2a1
  • ******O1a2a1a
  • ***O1a3
  • **O1b
  • ***O1b1
  • ****O1b1a
  • *****O1b1a1
  • ******O1b1a1a
  • *******O1b1a1a1
  • ********O1b1a1a1a
  • *********O1b1a1a1a1
  • **********O1b1a1a1a1a
  • ***********O1b1a1a1a1a1
  • ************O1b1a1a1a1a1a
  • *************O1b1a1a1a1a1a1
  • **************O1b1a1a1a1a1a1a
  • ***************O1b1a1a1a1a1a1a1
  • ****************O1b1a1a1a1a1a1a1a
  • *****************O1b1a1a1a1a1a1a1a1
  • ***************O1b1a1a1a1a1a1a2
  • ***************O1b1a1a1a1a1a1a3
  • **************O1b1a1a1a1a1a1b
  • ***************O1b1a1a1a1a1a1b1
  • *************O1b1a1a1a1a1a2
  • ***********O1b1a1a1a1a2
  • ************O1b1a1a1a1a2a
  • *************O1b1a1a1a1a2a1
  • ************O1b1a1a1a1a2b
  • **********O1b1a1a1a1b
  • ***********O1b1a1a1a1b1
  • ************O1b1a1a1a1b1a
  • *************O1b1a1a1a1b1a1
  • **************O1b1a1a1a1b1a1a
  • ***************O1b1a1a1a1b1a1a1
  • ************O1b1a1a1a1b1b
  • *************O1b1a1a1a1b1b1
  • ***********O1b1a1a1a1b2
  • ************O1b1a1a1a1b2a
  • *************O1b1a1a1a1b2a1
  • *********O1b1a1a1a2
  • **********O1b1a1a1a2a
  • ********O1b1a1a1b
  • *********O1b1a1a1b1
  • **********O1b1a1a1b1a
  • ***********O1b1a1a1b1a1
  • ***********O1b1a1a1b1a2
  • **********O1b1a1a1b1b
  • **********O1b1a1a1b1c
  • *********O1b1a1a1b2
  • *******O1b1a1a2
  • *******O1b1a1a3
  • ******O1b1a1b
  • *******O1b1a1b1
  • *****O1b1a2
  • ******O1b1a2a
  • *******O1b1a2a1
  • ********O1b1a2a1a
  • ******O1b1a2b
  • *******O1b1a2b1
  • ********O1b1a2b1a
  • *******O1b1a2b2
  • ******O1b1a2c
  • ***O1b2
  • ****O1b2a
  • *****O1b2a1
  • ******O1b2a1a
  • *******O1b2a1a1
  • ********O1b2a1a1a
  • *********O1b2a1a1a1
  • ********O1b2a1a1b
  • ********O1b2a1a1c
  • *******O1b2a1a2
  • ********O1b2a1a2a
  • *********O1b2a1a2a1
  • **********O1b2a1a2a1a
  • **********O1b2a1a2a1b
  • ***********O1b2a1a2a1b1
  • ********O1b2a1a2b
  • *******O1b2a1a3
  • ********O1b2a1a3a
  • ******O1b2a1b
  • *****O1b2a2
  • *O2
  • **O2a
  • ***O2a1
  • ****O2a1a
  • *****O2a1a1
  • ******O2a1a1a
  • *******O2a1a1a1
  • ********O2a1a1a1a
  • *********O2a1a1a1a1
  • **********O2a1a1a1a1a
  • ***********O2a1a1a1a1a1
  • ********O2a1a1a1b
  • *********O2a1a1a1b1
  • ********O2a1a1a1c
  • ******O2a1a1b
  • *******O2a1a1b1
  • ********O2a1a1b1a
  • *********O2a1a1b1a1
  • *********O2a1a1b1a1a
  • *********O2a1a1b1a2
  • ****O2a1b
  • ****O2a1c
  • *****O2a1c1
  • ******O2a1c1a
  • *******O2a1c1a1
  • ********O2a1c1a1a
  • *********O2a1c1a1a1
  • **********O2a1c1a1a1a
  • ***********O2a1c1a1a1a1
  • ************O2a1c1a1a1a1a
  • *************O2a1c1a1a1a1a1
  • **************O2a1c1a1a1a1a1a
  • ***************O2a1c1a1a1a1a1a1
  • ***************O2a1c1a1a1a1a1a2
  • *************O2a1c1a1a1a1a2
  • ************O2a1c1a1a1a1b
  • ***********O2a1c1a1a1a2
  • ************O2a1c1a1a1a2a
  • *********O2a1c1a1a2
  • *********O2a1c1a1a3
  • *********O2a1c1a1a4
  • **********O2a1c1a1a4a
  • *********O2a1c1a1a5
  • **********O2a1c1a1a5a
  • ***********O2a1c1a1a5a1
  • ************O2a1c1a1a5a1a
  • *************O2a1c1a1a5a1a1
  • **********O2a1c1a1a5b
  • ***********O2a1c1a1a5b1
  • *********O2a1c1a1a6
  • **********O2a1c1a1a6a
  • ***********O2a1c1a1a6a1
  • ***********O2a1c1a1a6a2
  • *********O2a1c1a1a7
  • *********O2a1c1a1a8
  • *********O2a1c1a1a9
  • ********O2a1c1a1b
  • ******O2a1c1b
  • *******O2a1c1b1
  • ********O2a1c1b1a
  • *********O2a1c1b1a1
  • *********O2a1c1b1a2
  • *******O2a1c1b2
  • ******O2a1c1c
  • *****O2a1c2
  • ***O2a2
  • ****O2a2a
  • *****O2a2a1
  • ******O2a2a1a
  • *******O2a2a1a1
  • ********O2a2a1a1a
  • *******O2a2a1a2
  • ********O2a2a1a2a
  • *********O2a2a1a2a1
  • **********O2a2a1a2a1a
  • ***********O2a2a1a2a1a1
  • ***********O2a2a1a2a1a2
  • ***********O2a2a1a2a1a3
  • *********O2a2a1a2a2
  • **********O2a2a1a2a2a
  • ***********O2a2a1a2a2a1
  • ********O2a2a1a2b
  • ******O2a2a1b
  • *****O2a2a2
  • ******O2a2a2a
  • *******O2a2a2a1
  • ****O2a2b
  • *****O2a2b1
  • ******O2a2b1a
  • *******O2a2b1a1
  • ********O2a2b1a1a
  • *********O2a2b1a1a1
  • **********O2a2b1a1a1a
  • ***********O2a2b1a1a1a1
  • ************O2a2b1a1a1a1a
  • *************O2a2b1a1a1a1a1
  • ***********O2a2b1a1a1a2
  • ************O2a2b1a1a1a2a
  • *************O2a2b1a1a1a2a1
  • **************O2a2b1a1a1a2a1a
  • ***************O2a2b1a1a1a2a1a1
  • *************O2a2b1a1a1a2a2
  • ***********O2a2b1a1a1a3
  • *********O2a2b1a1a2
  • **********O2a2b1a1a2a
  • ***********O2a2b1a1a2a1
  • ************O2a2b1a1a2a1a
  • *********O2a2b1a1a3
  • **********O2a2b1a1a3a
  • ***********O2a2b1a1a3a1
  • ***********O2a2b1a1a3a2
  • **********O2a2b1a1a3b
  • *********O2a2b1a1a4
  • **********O2a2b1a1a4a
  • ***********O2a2b1a1a4a1
  • ************O2a2b1a1a4a1a
  • ***********O2a2b1a1a4a2
  • *********O2a2b1a1a5
  • **********O2a2b1a1a5a
  • ***********O2a2b1a1a5a1
  • ***********O2a2b1a1a5a2
  • **********O2a2b1a1a5b
  • ***********O2a2b1a1a5b1
  • *********O2a2b1a1a6
  • **********O2a2b1a1a6a
  • **********O2a2b1a1a6b
  • ***********O2a2b1a1a6b1
  • ************O2a2b1a1a6b1a
  • ************O2a2b1a1a6b1b
  • ***********O2a2b1a1a6b2
  • ***********O2a2b1a1a6b3
  • *********O2a2b1a1a7
  • **********O2a2b1a1a7a
  • ***********O2a2b1a1a7a1
  • **********O2a2b1a1a7b
  • *********O2a2b1a1a8
  • **********O2a2b1a1a8a
  • ********O2a2b1a1b
  • *******O2a2b1a2
  • ********O2a2b1a2a
  • *********O2a2b1a2a1
  • **********O2a2b1a2a1a
  • ***********O2a2b1a2a1a1
  • ************O2a2b1a2a1a1a
  • *************O2a2b1a2a1a1a1
  • ************O2a2b1a2a1a1b
  • ***********O2a2b1a2a1a2
  • ************O2a2b1a2a1a2a
  • *************O2a2b1a2a1a2a1
  • **************O2a2b1a2a1a2a1a
  • ***********O2a2b1a2a1a3
  • ************O2a2b1a2a1a3a
  • *************O2a2b1a2a1a3a1
  • ************O2a2b1a2a1a3b
  • *************O2a2b1a2a1a3b1
  • *************O2a2b1a2a1a3b2
  • **************O2a2b1a2a1a3b2a
  • ***************O2a2b1a2a1a3b2a1
  • **************O2a2b1a2a1a3b2b
  • ***************O2a2b1a2a1a3b2b1
  • ****************O2a2b1a2a1a3b2b1a
  • ***************O2a2b1a2a1a3b2b2
  • **********O2a2b1a2a1b
  • ***********O2a2b1a2a1b1
  • ************O2a2b1a2a1b1a
  • **********O2a2b1a2a1c
  • **********O2a2b1a2a1d
  • ***********O2a2b1a2a1d1
  • ************O2a2b1a2a1d1a
  • ************O2a2b1a2a1d1b
  • ********O2a2b1a2b
  • *********O2a2b1a2b1
  • **********O2a2b1a2b1a
  • ***********O2a2b1a2b1a1
  • ***********O2a2b1a2b1a2
  • *********O2a2b1a2b2
  • **********O2a2b1a2b2a
  • ********O2a2b1a2c
  • *****O2a2b2
  • ******O2a2b2a
  • *******O2a2b2a1
  • ********O2a2b2a1a
  • *********O2a2b2a1a1
  • *********O2a2b2a1a2
  • ********O2a2b2a1b
  • *********O2a2b2a1b1
  • *********O2a2b2a1b2
  • **********O2a2b2a1b2a
  • *******O2a2b2a2
  • ********O2a2b2a2a
  • *********O2a2b2a2a1
  • *********O2a2b2a2a2
  • ********O2a2b2a2b
  • *********O2a2b2a2b1
  • *********O2a2b2a2b2
  • **********O2a2b2a2b2a
  • **********O2a2b2a2b2b
  • ***********O2a2b2a2b2b1
  • **********O2a2b2a2b2c
  • ******O2a2b2b
  • *******O2a2b2b1
  • ********O2a2b2b1a
  • *********O2a2b2b1a1
  • ********O2a2b2b1b
  • ***O2a3
  • ***O2a4
  • **O2b
  • ***O2b1
  • ****O2b1a
  • *****O2b1a1
  • ***O2b2
  • ****O2b2a