Berothidae


The Berothidae are a family of winged insects of the order Neuroptera. They are known commonly as the beaded lacewings. The family was first named by Anton Handlirsch in 1906. The family consists of 24 genera and 110 living species distributed discontinuously worldwide, mostly in tropical and subtropical regions. Numerous extinct species have also been described. Their ecology is poorly known, but in the species where larval stages have been documented, the larvae are predators of termites.

Systematics

A considerable fossil diversity of beaded lacewings is known from the Late Jurassic onwards, containing numerous genera which are likewise basal or incertae sedis.
;Subfamily Berothimerobiinae
  • Berothimerobius
;Subfamily Berothinae
;Subfamily Cyrenoberothinae
  • †Aggregataberotha
  • †Araripeberotha Cretaceous ; Crato Formation, Brazil
  • †Caririberotha Cretaceous ; Crato Formation, Brazil
  • Cyrenoberotha
  • Manselliberotha
  • †Microberotha
  • Ormiscocerus
  • †Protoberotha
  • †Sibelliberotha
  • Speleoberotha
;Subfamily †Mesithoninae
;Subfamily Nosybinae
  • †Banoberotha Cretaceous ; Lebanese amber, Lebanon
  • Naizema
  • Nosybus
  • Tanzanberotha
;Subfamily Nyrminae
  • Nyrma
;Subfamily Protobiellinae
  • Austroberothella
  • Protobiella
;Subfamily Trichomatinae
  • Trichoberotha
  • Trichoma
;Subfamily Incertae sedis
  • †Ansoberotha Cenomanian; Burmese amber, Myanmar
  • †Berothone Callovian/Oxfordian; Karabastau Formation, Kazakhstan
  • †Cantabroberotha Cretaceous ; Spanish amber, Spain
  • †Cornoberotha Cenomanian; Burmese amber, Myanmar
  • †Dasyberotha Cenomanian; Burmese amber, Myanmar
  • †Epimesoberotha Cretaceous ; Lulworth Formation, United Kingdom
  • †Ethiroberotha Cenomanian; Burmese amber, Myanmar
  • †Haploberotha Cenomanian; Burmese amber, Myanmar
  • †Iceloberotha Cenomanian; Burmese amber, Myanmar
  • †Jersiberotha Cenomanian; Burmese amber, Myanmar; Cretaceous ; New Jersey amber, United States
  • †Krokhathone Jurassic ; Karabastau Formation, Kazakhstan
  • †Maculaberotha Cenomanian; Burmese amber, Myanmar
  • †Magniberotha Cenomanian; Burmese amber, Myanmar
  • †Nascimberotha Turonian; New Jersey amber, United States
  • †Plesiorobius Cretaceous ; Canadian amber, Canada; Campanian; Taymyr amber, Russia; Cenomanian, unknown formation, Russia
  • †Proberotha Lutetian, Baltic Amber, northern Europe
  • †Osmyloberotha Cenomanian; Burmese amber, Myanmar
  • †Sinosmylites Daohugou, Haifanggou Formation, China, Callovian, Karabastau Formation, Kazakhstan, Callovian/Oxfordian, Ulaan-Ereg Formation, Mongolia, Late Jurassic
  • †Systenoberotha Cenomanian; Burmese amber, Myanmar
  • †Telistoberotha Cenomanian; Burmese amber, Myanmar
  • †Xiaoberotha Cenomanian; Burmese amber, Myanmar

    Formerly included taxa

Subfamily Paraberothinae
  • †Acanthoberotha Cenomanian; Burmese amber, Myanmar
  • †Alboberotha Cretaceous ; Charentense amber, France
  • †Astioberotha Cenomanian; Burmese amber, Myanmar
  • †Chimerhachiberotha Barremian; Lebanese amber, Lebanon
  • †Creagroparaberotha Cenomanian; Burmese amber, Myanmar
  • †Eorhachiberotha Cenomanian; Burmese amber, Myanmar
  • †Kujiberotha Cretaceous ; Kuji amber, Japan
  • †Micromantispa Cenomanian; Burmese amber, Myanmar
  • †Paraberotha Barremian; Lebanese amber, Lebanon
  • †Pseudosisyra Aptian; Zaza Formation, Russia
  • †Raptorapax Barremian; Lebanese amber, Lebanon
  • †Retinoberotha Cenomanian; Bezonnais amber, France,
  • †Rhachibermissa Turonian; New Jersey amber, United States
  • †Scoloberotha Cenomanian; Burmese amber, Myanmar
  • †Spinoberotha Cenomanian; Burmese amber, Myanmar
  • †Stygioberotha Cenomanian; Burmese amber, Myanmar
  • †Uranoberotha Cenomanian; Burmese amber, Myanmar