List of Mesozoic birds


This list of Mesozoic birds is a comprehensive list of all Mesozoic dinosaurs that have been assigned to the clade Avialae. The list includes all commonly accepted genera, but also genera that are now considered invalid, doubtful, or were not formally published, as well as junior synonyms of more established names, and genera that are no longer considered avialan. The list currently includes genera.

Scope and terminology

There is no official, canonical list of Mesozoic bird genera, but some of the more complete ones include Holtz's list of Mesozoic dinosaurs, Molina-Perez & Larramendi's list of theropods, and Mickey Mortimer's Theropod Database. Lists of ex-birds are generally rare, but one example is the "Ex-Saurischian Taxa and Basal Saurischians" section of the Theropod Database.

Authors and year

The authors column lists the authors of the formal description responsible for the erection of the genus listed. They are not necessarily the same as the authors of the type species, as sometimes a species from one genus is determined sufficiently distinct to warrant the erection of a new genus to house it. If this is the case, only the latter authors will be listed. The year column notes the year the genus' description was published.

Status

Naming conventions and terminology follow the International Code of Zoological Nomenclature. Technical terms used include:Junior synonym: A name which describes the same taxon as a previously published name. If two or more genera are formally designated and the type specimens are later assigned to the same genus, the first to be published is the senior synonym, and all other instances are junior synonyms. Senior synonyms are generally used, except by special decision of the ICZN, but junior synonyms cannot be used again, even if deprecated. Junior synonymy is often subjective, unless the genera described were both based on the same type specimen.Nomen nudum : A name that has appeared in print but has not yet been formally published by the standards of the ICZN. Nomina nuda are invalid, and are therefore not italicized as a proper generic name would be. If the name is later formally published, that name is no longer a nomen nudum and will be italicized on this list. Often, the formally published name will differ from any nomina nuda that describe the same specimen.Preoccupied name: A name that is formally published, but which has already been used for another taxon. This second use is invalid and the name must be replaced. Nomen dubium : A name describing a fossil with no unique diagnostic features. As this can be an extremely subjective and controversial designation, this term is not used on this list.

Age

The age column denotes the epoch of geologic time to which the fossils date. Genera that are invalid, misidentified, or otherwise do not represent a valid Mesozoic bird are listed as age N/A because there was never a time in which a Mesozoic bird by that generic name actually lived.

Location and notes

The location column designates the geographic region where remains of the relevant genus have been found. The regions used are countries, despite the fact that there were no political boundaries in Mesozoic times; they are only used for convenience. Genera that are invalid, misidentified, or otherwise do not represent a valid Mesozoic bird are listed as location N/A because there was never a place in which a Mesozoic bird by that generic name actually lived. The notes column is a collection of annotations on the scientific significance and taxonomic history of listed genera, as well as elaborations on the information presented in other columns.

List

GenusAuthorsYearStatusAgeLocationNotesImages
AbavornisPanteleyev1998ValidUzbekistan
AberratiodontusGong et al.2004Junior synonymChinaJunior synonym of Yanornis.
AbitusavisWang et al.2020ValidChina
AlamitornisAgonlin & Martinelli2009ValidArgentina
AlcmonavisRauhut et al.2019ValidGermany
AlethoalaornisLi et al.2007ValidChina
AlexornisBrodkorb1976ValidMexico
AmbiortusKurochkin1982ValidMongolia
AmbopteryxWang et al.2019ValidChinaConsidered a non-avialan theropod by some researchers.
AnatalavisOlson & Parris1987ValidUnited States
England
The type species is known from the Late Cretaceous–Paleocene border, while a referred species is from the Eocene.
AnchiornisXu et al.2009ValidChinaConsidered a non-avialan theropod by some researchers.
AngelinornisKashin1972Junior synonymUnited StatesJunior synonym of Ichthyornis.
AntarcticavisCordes-Person et al.2020ValidAntarctica
ApatornisMarsh1873ValidUnited States
ApsaravisNorell & Clarke2001ValidMongolia
ArchaeopteryxMeyer1861ValidGermanyConsidered a non-avialan theropod by some researchers.
ArchaeorhynchusZhou & Zhang2006ValidChina
ArchaeornisPetronievics1917Junior synonymGermanyJunior synonym of Archaeopteryx.
ArchaeornithuraWang et al.2015ValidChina
ArchaeovolansCzerkas & Xu2002Junior synonymChinaJunior synonym of Yanornis.
AsiahesperornisNesov & Prizemlin1991ValidKazakhstan
AsteriornisField et al.2020ValidBelgium
AurornisGodefroit et al.2013ValidChinaConsidered a non-avialan theropod by some researchers.
AustinornisClarke2004ValidUnited States
AvimaiaBailleul et al.2019ValidChina
AvisaurusBrett-Surman & Paul1985ValidUnited States
BalaurCsiki et al.2010ValidRomaniaOriginally thought to be a non-avialan theropod.
BaminornisChen et al.2025ValidChina-
BaptornisMarsh1877ValidUnited States
BauxitornisDyke & Ősi2010ValidHungary
BeiguornisWang et al.2022ValidChina
BelluliaWang et al.2016aPreoccupiedN/AN/AName preoccupied by the moth Bellulia Fibiger 2008 and replaced by Bellulornis.
BellulornisWang et al.2016bValidChina
BohaiornisHu et al.2011ValidChina
BoluochiaZhou1995ValidChina
BradycnemeHarrison & Walker1975MisidentificationN/AN/AOriginally described as a bird; more likely a non-avialan theropod.
BrevidentavisO'Connor et al.2021ValidChina
BrevirostruavisLi et al.2021ValidChina
BrodavisMartin et al.2012ValidUnited States
Mongolia
CaihongHu et al.2018ValidChinaConsidered to be a non-avialan theropod by some researchers.
CamptodontornisDemirjian2019ValidChina
CamptodontusLi et al.2010PreoccupiedN/AN/AName preoccupied by the beetle Camptodontus Dejean, 1826 and replaced by Camptodontornis.
CanadagaHou1999ValidCanada
CastignovolucrisBuffetaut et al.2023ValidFrance-
CatenoleimusPanteleyev1998ValidUzbekistan
CathayornisZhou et al.1992ValidChina
CeramornisBrodkorb1963ValidUnited States
CerebavisKurochkin et al.2006ValidRussia
ChangchengornisJi et al.1999ValidChina
ChangmaornisWang et al.2013ValidChina
ChangzuiornisHuang et al.2016DisputedChinaPossibly synonymous with Juehuaornis.
ChaoyangiaHou & Zhang1993ValidChina
ChiappeavisO'Connor et al.2016ValidChina
ChongmingiaWang et al.2016ValidChina
ChromeornisO'Connor et al.2025ValidChina
ChupkaornisTanaka et al.2017ValidJapan
CimoliornisOwen1846MisidentificationN/AN/AOriginally described as a bird; actually a pterosaur.
CimolopteryxMarsh1892ValidUnited StatesMay be Paleocene.
ColonosaurusMarsh1872Junior synonymUnited StatesJunior synonym of Ichthyornis.
ConcornisSanz & Buscalioni1992ValidSpain
ConfuciusornisHou et al.1995ValidChina
ConiornisMarsh1893Junior synonymUnited StatesJunior synonym of Hesperornis.
CosesaurusEllenberger & de Villalta1974MisidentificationN/AN/ASometimes proposed to be a bird ancestor; actually a tanystropheid.
CratoavisCarvalho et al.2015ValidBrazil
CratonavisLi et al.2023ValidChina
CretaaviculusBazhanov1969ValidKazakhstan
CretornisFritsch1880MisidentificationN/AN/AOriginally described as a bird; actually a pterosaur.
CruralispenniaWang et al.2017ValidChina
CuspirostrisornisHou1997ValidChina
DalianraptorGao & Liu2005DisputedN/AN/ALikely a chimaera consisting of a Jeholornis and another theropod.
DalingheornisZhang et al.2006ValidChina
DapingfangornisLi et al.2006ValidChina
DidactylornisYuan2008Junior synonymChinaJunior synonym of Sapeornis.
DingavisO'Connor et al.2016DisputedChinaPossibly synonymous with Changzuiornis and/or Juehuaornis.
DunhuangiaWang et al.2015ValidChina
ElbretornisWalker & Dyke2009ValidArgentinaPossibly synonymous with Lectavis, Soroavisaurus and/or Yungavolucris.
ElektorornisXing et al.2019ValidMyanmar
ElopteryxAndrews1913MisidentificationN/AN/AOriginally described as a bird; more likely to be a non-avialan theropod.
ElsornisChiappe et al.2007ValidMongolia
EnaliornisSeeley1876ValidEngland
EnantiophoenixCau & Arduini2008ValidLebanon
EnantiornisWalker1981ValidArgentina
EoalulavisSanz et al.1996ValidSpain
EocathayornisZhou2002ValidChina
EoconfuciusornisZhang et al.2008ValidChina
EoenantiornisHou et al.1999ValidChina
EogranivoraZheng et al.2018ValidChina
EopengornisWang et al.2014ValidChina
EosinopteryxGodefroit et al.2013ValidChinaConsidered a non-avialan theropod by some researchers.
EpidendrosaurusZhang et al.2002Junior synonymChinaPossible synonym of Scansoriopteryx.
EpidexipteryxZhang et al.2008ValidChinaConsidered a non-avialan theropod by some researchers.
EurolimnornisKessler & Jurcsák1986MisidentificationN/AN/AOriginally described as a bird; reinterpreted as a pterodactyloid pterosaur.
EvgenavisO'Connor et al.2014ValidRussia
ExplorornisPanteleyev1998ValidUzbekistan
FalcatakelyO'Connor et al.2020ValidMadagascar
FeitianiusO'Connor et al.2015ValidChina
FlexomornisTykoski & Fiorillo2010ValidUnited States
FortipesavisClark & O'Connor2021ValidMyanmar
FortunguavisWang et al.2014ValidChina
FujianvenatorXu et al.2023ValidChinaFound within the family Anchiornithidae, which may represent non-avialan dinosaurs
FukuipteryxImai et al.2019ValidJapan
FumicollisBell & Chiappe2015ValidUnited States
GallornisLambrecht1931ValidFrance
GansusHou & Liu1984ValidChina
GargantuavisBuffetaut & Le Loeuff1998ValidFrance
Spain
Romania
GettyiaAtterholt et al.2018ValidUnited States
GobipipusKurochkin & Chatterjee2013ValidMongoliaVery similar to embryonic Gobipteryx.
GobipteryxElżanowski1994ValidMongolia
GrabauornisDalsätt et al.2014ValidChina
GracilornisLi & Hou2011ValidChina
GraculavusMarsh1872ValidUnited StatesMay be Paleocene.
GretcheniaoChiappe et al.2019ValidChina
GriphornisWoodward1862Junior synonymGermanyName officially rejected in favor of Archaeopteryx.
GriphosaurusWagner1862Junior synonymGermanyName officially rejected in favor of Archaeopteryx.
GuildavisClarke2004ValidUnited States
GurilyniaKurochkin1999ValidMongolia
HalimornisChiappe2002ValidUnited States
HargeriaLucas1903Junior synonymUnited StatesJunior synonym of Hesperornis.
"Hebeiornis"Xu et al.1999Nomen nudumN/AN/ASynonym of Vescornis due to being based on the same holotype. However, despite predating the description of Vescornis by five years, its description is so poor that it is regarded as a nomen nudum and a synonym.
HeptasteornisHarrison & Walker1975MisidentificationN/AN/AOriginally described as a bird; more likely a non-avialan theropod.
HesperornisMarsh1872ValidUnited States
Canada
Russia
HolbotiaZelenkov & Averianov2015ValidMongolia
HollandaBell et al.2010ValidMongolia
HongshanornisZhou & Zhang2005ValidChina
HorezmavisNessov & Borkin1983ValidUzbekistan
HouornisWang & Liu2015ValidChina
HuoshanornisWang et al.2010ValidChina
IaceornisClarke2004ValidUnited States
IberomesornisSanz & Bonaparte1992ValidSpain
IchthyornisMarsh1873ValidUnited States
IlerdopteryxLacasa-Ruiz1985ValidSpain
ImparavisWang et al.2024ValidChina
IncolornisPanteleyev1998ValidUzbekistan
IntiornisNovas et al.2010ValidArgentina
IteravisZhou et al.2014ValidChina
JanavisBenito et al.2022ValidBelgium-
JeholornisZhou et al.2002ValidChina
JianchangornisZhou et al.2009ValidChina
JibeiniaHou1997ValidChinaOnly known specimen missing.
JinfengopteryxJi et al.2005MisidentificationN/AN/AOriginally described as a bird; actually a non-avian dinosaur.
JinguofortisWang et al.2018ValidChina
JinzhouornisHou et al.2002Junior synonymChinaJunior synonym of Confuciusornis.
JiuquanornisWang et al.2013ValidChina
JixiangornisJi et al.2002ValidChina
JudinornisNessov & Borkin1983ValidMongolia
JuehuaornisWang et al.2015ValidChina
JunornisLiu et al.2017ValidChina
JurapteryxHowgate1984Junior synonymGermanyJunior synonym of Archaeopteryx.
KaririavisCarvalho et al.2021ValidBrazil
KhinganornisWang et al.2020ValidChina
KizylkumavisNessov1984ValidUzbekistan
KompsornisWang et al.2020ValidChina
KookneNovas et al.2019ValidArgentina
KunpengornisHuang et al.2025ValidChina
KuszholiaNesov1992ValidUzbekistan
LamarqueavisAgnolin2010ValidArgentina
United States
Canada
LaopteryxMarsh1881MisidentificationN/AN/AOriginally described as a bird; actually a pterosaur.
LaornisMarsh1870ValidUnited StatesMay be Paleocene.
LargirostrornisHou1997DisputedChinaPossible synonym of Cathayornis.
LectavisChiappe1993ValidArgentina
LenesornisKurochkin1996ValidUzbekistan
LestornisMarsh1876Junior synonymUnited StatesJunior synonym of Hesperornis.
LiaoningornisHou1996ValidChina
LiaoxiornisHou & Chen1999ValidChina
LimenavisClarke & Chiappe2001ValidArgentina
LimnornisKessler & Jurcsak1984MisidentificationN/AN/AName preoccupied by the ovenbird Limnornis Gould 1839 and replaced by Palaeocursornis. Originally described as a bird; actually a pterosaur.
LingyuanornisJi & Ji1999Junior synonymN/AN/AJunior synonym of Liaoxiornis as it is based on the same holotype.
LinyiornisWang et al.2016ValidChina
LisboasaurusSeiffert1970MisidentificationN/AN/ASometimes proposed to be a bird; actually a crocodilian.
LonchodytesBrodkorb1963ValidUnited StatesMay be Paleocene.
LongchengornisHou1997DisputedChinaPossible synonym of Cathayornis.
LongicrusavisO'Connor et al.2010ValidChina
LongipteryxZhang et al.2001ValidChina
LongirostravisHou et al.2004ValidChina
LongisquamaSharov1970MisidentificationN/AN/ASometimes proposed to be a bird ancestor; actually an indeterminate reptile.
LongusunguisWang et al.2014ValidChina
MaaqwiMcLachlan et al.2017ValidCanada
MagnusavisClark et al.2024ValidUnited States
MartinavisWalker et al.2007ValidFrance
Argentina
MeemannavisO'Connor et al.2021ValidChina
MegalancosaurusCalzavara et al.1980MisidentificationN/AN/ASometimes proposed to be a bird ancestor, but actually a drepanosaurid.
MengciusornisWang et al.2019ValidChina
MicroenantiornisWei & Li2017ValidChina
MirarceAtterholt et al.2018ValidUnited States
MirusavisWang et al.2020ValidChina
MonoenantiornisHu & O'Connor2016ValidChina
MusivavisWang et al.2022ValidChina
MystiornisKurochkin et al.2011ValidRussia
NanantiusMolnar1986ValidAustralia
NavaornisChiappe et al.2024ValidBrazil
NeimengornisWang et al.2021ValidChina
NeobohaiornisShen et al.2024ValidChina
NeogaeornisLambrecht1929ValidChile
NeuquenornisChiappe & Calvo1994ValidArgentina
NoguerornisLacasa-Ruiz1989ValidSpain
NovacaesarealaParris & Hope2002ValidUnited StatesMay be Paleocene.
NovavisO'Connor et al.2025In pressChina
OculudentavisXing et al.2020MisidentificationN/AN/AOriginally described as a bird; reinterpreted as an indeterminate lizard.
OmnivoropteryxCzerkas & Ji2002ValidChina
OrienantiusLiu et al.2019ValidChina
OstromiaFoth & Rauhut2017ValidGermanyConsidered a non-avialan theropod by some researchers.
OtogornisHou1994ValidChina
PalaeocursornisKessler & Jurcsak1986MisidentificationN/AN/AOriginally described as a bird; actually a pterosaur.
PalaeonornisEmmons1857MisidentificationN/AN/AOriginally described as a bird; actually a junior synonym of the phytosaur Rutiodon.
PalaeopteryxJensen1981MisidentificationN/AN/AOriginally described as a bird; more likely a non-avian theropod.
PalaeornisMantell1844MisidentificationN/AN/AName preoccupied by the parrot Palaeornis Vigors 1825. Originally described as a bird; actually a pterosaur.
PalaeotringaMarsh1870ValidUnited StatesMay be Paleocene.
PalintropusBrodkorb1970ValidUnited States
Canada
ParabohaiornisWang et al.2014ValidChina
ParahesperornisMartin1984ValidUnited States
ParahongshanornisLi et al.2011ValidChina
ParapengornisHu et al.2015ValidChina
ParaprotopteryxZheng et al.2007ValidChina
ParascaniornisLambrecht1933Junior synonymSwedenJunior synonym of Baptornis.
ParvavisWang et al.2014ValidChina
PasquiaornisTokaryk et al.1997ValidCanada
PatagopteryxAlvarenga & Bonaparte1992ValidArgentina
PedopennaXu & Zhang2005ValidChinaConsidered a non-avialan theropod by some researchers.
PelagornisSeeley1866PreoccupiedN/AN/ANamed preoccupied by the Miocene bird Pelagornis Lartet 1857 and replaced by Enaliornis.
PengornisZhou et al.2008ValidChina
PiksiVarricchio2002MisidentificationN/AN/AOriginally described as a bird; reinterpreted as a pterodactyloid pterosaur.
PiscivoravisZhou et al.2013ValidChina
PiscivorenantiornisWang & Zhou2017ValidChina
PlatanavisNesov1992ValidUzbekistan
PlegadornisWetmore1962PreoccupiedN/AN/AName preoccupied by the modern bird Plegadornis Brehm 1855; junior synonym of Ichthyornis.
PolarornisChatterjee2002ValidAntarctica
PotamornisElzanowski et al.2001ValidUnited States
PraeornisRautian1978ValidKazakhstan
PriscavolucrisGomez Pallerola1979MisidentificationN/AN/AOriginally described as a bird; reinterpreted as a hybodontiform fish.
"Proornis"Lim1993Nomen nudumN/AN/A
ProtarchaeopteryxJi & Ji1997MisidentificationN/AN/AOriginally described as a bird; actually a non-avian dinosaur.
ProtoavisChatterjee1991MisidentificationN/AN/AOriginally described as a bird; more likely a chimaera of several vertebrates.
ProtopteryxZhang & Zhou2000ValidChina
PterodactylusRafinesque1815MisidentificationN/AN/ASometimes thought to be a bird before being described as a pterosaur.
PterygornisWang et al.2015ValidChina
PujatopouliIrazoqui et al.2025ValidAntarctica
QilianiaJi et al.2011ValidChina
RahonaForster et al.1998aPreoccupiedN/AN/AName preoccupied by the moth Rahona Griveaud 1975 and replaced by Rahonavis.
RahonavisForster et al.1998bValidMadagascarConsidered a non-avialan theropod by some researchers.
RapaxavisMorschhauser et al.2009ValidChina
RhamphorhynchusMeyer1846MisidentificationN/AN/AOriginally thought to be a bird before being described as a pterosaur.
SamrukiaNaish et al.2012MisidentificationN/AN/AOriginally described as a bird; actually a pterosaur.
SapeornisZhou & Zhang2002ValidChina
SazavisNesov1989ValidUzbekistan
ScansoriopteryxCzerkas & Yuan2002ValidChinaConsidered a non-avialan theropod by some researchers.
SchizoouraZhou et al.2012ValidChina
SerikornisLefèvre et al.2017ValidChinaConsidered a non-avialan theropod by some researchers.
ShangyangWang & Zhou2019ValidChina
ShanweiniaoO'Connor et al.2009ValidChina
ShengjingornisLi et al.2012ValidChina
ShenqiornisWang et al.2010ValidChina
ShenshiornisHu2010Junior synonymChinaJunior synonym of Sapeornis.
ShenzhouraptorJi et al.2002DisputedChinaPossible synonym of Jeholornis.
ShuilingornisWang et al.2025ValidChina
SimiliyanornisWang et al.2020ValidChina
SinornisSereno & Rao1992ValidChina
SonglingornisHou1997ValidChina
SoroavisaurusChiappe1993ValidArgentina
StruthiosaurusBunzel1871MisidentificationN/AN/AOriginally thought to be a bird; actually an ankylosaurian dinosaur.
"Styginetta"Stidham2001Nomen nudumN/AN/AObscure; named in a thesis.
SulcavisO'Connor et al.2013ValidChina
TelmatornisMarsh1870ValidUnited States
TeviornisKurochkin et al.2002ValidMongolia
TianyuornisZheng et al.2014ValidChina
TingmiatornisBono et al.2016ValidCanada
TorotixBrodkorb1963ValidUnited States
TytthostonyxOlson & Parris1987ValidUnited States
VegavisClarke et al.2005ValidAntarctica
VescornisZhang et al.2004DisputedChinaPossible junior synonym of Jibeinia.
VolgavisNessov & Yarkov1989ValidRussia
VoronaForster et al.1996ValidMadagascar
WasaibpanchiMalkani2021DisputedPakistanPublication status uncertain.
WellnhoferiaElżanowski2001DisputedGermanyPossible junior synonym of Archaeopteryx.
WyleyiaHarrison & Walker1973ValidEngland
XiangornisHu et al.2012ValidChina
XinghaiornisWang et al.2013ValidChina
XiaotingiaXu et al.2011ValidChinaConsidered a non-avialan theropod by some researchers.
YandangornisCai & Zhao1999ValidChina
YangavisWang et al.2018ValidChina
YanornisZhou & Zhang2001ValidChina
YatenavisHerrera et al.2022ValidArgentina
YiXu et al.2015ValidChinaConsidered a non-avialan theropod by some researchers.
YixianornisZhou & Zhang2001ValidChina
YixianosaurusXu & Wang2003ValidChinaConsidered a non-avialan theropod by some researchers.
YuanchuavisWang et al.2021ValidChina
YuanjiawaornisHu et al.2015ValidChina
YumenornisWang et al.2013ValidChina
YungavolucrisChiappe1993ValidArgentina
YuornisXu et al.2021ValidChina
ZhongjianornisZhou et al.2010ValidChina
ZhongornisGao et al.2008ValidChina
ZhouornisZhang et al.2013ValidChina
ZhylgaiaNessov1988ValidKazakhstanDating is uncertain; may be Late Cretaceous or Paleogene.
ZhyraornisNessov1984ValidUzbekistan