Metriorhynchidae


Metriorhynchidae is an extinct family of specialized, aquatic metriorhynchoid crocodyliforms from the Middle Jurassic to the Early Cretaceous period of Europe, North America and South America. The name Metriorhynchidae was coined by the Austrian zoologist Leopold Fitzinger in 1843. The group contains two subfamilies, the Metriorhynchinae and the Geosaurinae. They represent the most marine adapted of all archosaurs.

Description

Metriorhynchids are fully aquatic crocodyliforms. Their forelimbs were small and paddle-like, and unlike living crocodylians, they lost their osteoderms. Their body shape maximised hydrodynamy, as they did have a shark-like tail fluke. Like ichthyosaurs and plesiosaurs, metriorhynchids developed smooth, scaleless skin. However, unlike those marine reptiles, they were poikilothermic with little to no endothermic ability.
Metriorhynchids were the only group of archosaurs to become fully adapted to the marine realm, becoming pelagic in lifestyle. With tail flukes, reduced limb musculature, and long bones histologically comparable to other obligately aquatic animals, they were almost certainly incapable of terrestrial locomotion; combined with an unusually tall hip opening, as also seen in other obligately aquatic reptiles including the viviparous Keichousaurus, these characters suggest that metriorhynchids gave live birth. A fossil of a pregnant Dakosaurus female recovered from the Late Jurassic plattenkalk, Bavaria, preserves the complete skeleton of a neonate with small, paddle-like forelimbs unsuited for walking on land, similar to those of adults, further supporting live birth in metriorhynchids. Recent research posits that despite their successful adaptation to a pelagic lifestyle, basal metriorhynchids were uniquely disadvantaged among aquatic tetrapods in evolving into sustained swimmers due to little to no posterodorsal retraction of the external nares. The family has a wide geographic distribution, with material found in Argentina, Chile, Cuba, England, France, Germany, Italy, Mexico, Poland, Russia, Switzerland and Czech Republic.

Classification

analyses published during the 2000s cast doubt on the idea that many traditional metriorhynchid genera formed natural groups. The traditional species of Geosaurus, Dakosaurus and Cricosaurus were found to represent unnatural groups, and the species traditionally classified in these genera were reshuffled in a study published in November 2009 by Mark T. Young and Marco Brandalise de Andrade. The monophyly of Metriorhynchus and Teleidosaurus is also unsupported, and the species of these genera are pending reclassification.
The classification presented by Young and Andrade in 2009 was approved in later studies of the Metriorhynchidae. Metriorhynchidae is a node-based taxon defined in the PhyloCode by Mark T. Young and colleagues in 2024 as "the smallest clade within Metriorhynchoidea containing Thalattosuchus superciliosus, Gracilineustes leedsi, Metriorhynchus brevirostris, Rhacheosaurus gracilis, and Geosaurus giganteus" The cladogram below follows the topology from the 2020 analyses by Young et al. and reduced to genera only.

List of genera

The type genus of the family Metriorhynchidae is Metriorhynchus from the Middle to Late Jurassic. Other genera included within this family are Cricosaurus, Geosaurus, and Dakosaurus. Though once considered a metriorhynchid, Teleidosaurus has since been found to be slightly more distantly related to these animals within the superfamily Metriorhynchoidea.
Within this family, the genus Neustosaurus and Enaliosuchus are considered nomen dubium.
The genus Capelliniosuchus was once thought to be a metriorhynchid similar to Dakosaurus. However, it was later found to be a mosasaur.
GenusStatusAuthorAgeLocationDescriptionSynonymsImages

  • Aggiosaurus
Ambayrac, 1913late OxfordianFrance.Sometimes considered to be a nomen dubium.
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|
Suchodus durobrivensis

  • Cricosaurus
ValidWagner, 1858middle Oxfordian to upper ValanginianArgentina, Chile, Cuba, England, France, Germany, Mexico, Russia and Switzerland.
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|
Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|
Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|
Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai

Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis

  • Dakosaurus
Validvon Quenstedt, 1856upper Kimmeridgian to lower BerriasianArgentina, England, France, Germany, Mexico, Poland, Switzerland and possibly Russia.
  • Dacosaurus
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|
Suchodus durobrivensis

  • Enaliosuchus
nomen dubiumKoken, 1883ValanginianGermany.-
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|
Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|
Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|
Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai

Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis

  • Geosaurus
ValidCuvier, 1824upper Kimmeridgian to upper ValanginianFrance, Germany, Italy and Switzerland.
  • Brachytaenius
  • Halilimnosaurus
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|
Suchodus durobrivensis

  • Gracilineustes
ValidYoung et al., 2010middle Callovian to lower KimmeridgianEngland and France.
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|
Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|
Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|
Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai

Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis

  • Maledictosuchus
ValidParrilla-Bel et al., 2013middle Callovian to KimmeridgianSpain and Mexico.
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|
Suchodus durobrivensis

  • Metriorhynchus
Validvon Meyer, 1832lower Kimmeridgian.France.
  • Metiorychus
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|
Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|
Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|
Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai

Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis

  • Neptunidraco
ValidCau & Fanti, 2011earliest BathonianItaly.
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|
Suchodus durobrivensis

  • Neustosaurus
nomen dubiumValanginian— potential senior synonym of EnaliosuchusImage:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|
Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|
Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|
Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai

Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis

  • Plesiosuchus
ValidOwen, 1884late Kimmeridgian to early TithonianEngland and possibly Spain.
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|
Suchodus durobrivensis

  • Purranisaurus
ValidRusconi, 1948upper TithonianArgentina and Chile.
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|
Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|
Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|
Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai

Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis

  • Rhacheosaurus
Validvon Meyer, 1831lower TithonianGermany.
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|
Suchodus durobrivensis

  • Suchodus
ValidLydekker, 1890middle Callovian to lower OxfordianEngland and France.
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|
Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|
Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|
Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai

Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis

  • Thalattosuchus
ValidYoung et al., 2020lower Callovian to lower OxfordianEngland and France.
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|
Suchodus durobrivensis

  • Torvoneustes
ValidAndrade et al., 2010Kimmeridgian to TithonianEngland.
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|
Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|
Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|
Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai

Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis

  • Tyrannoneustes
ValidYoung et al., 2013CallovianEngland.
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|
Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|
Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|
Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|
Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai

Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|
Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|
Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|
Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|
Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai

Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|
Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|
Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|
Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|
Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai

Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|
Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|
Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|
Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|
Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai

Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|
Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|
Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|
Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|
Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai

Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|
Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|
Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|
Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|
Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai

Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|
Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|
Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|
Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|
Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai

Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|
Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|
Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|
Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|
Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai

Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|
Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|
Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|
Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|
Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai

Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|
Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|
Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|
Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|
Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai

Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|
Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|
Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|
Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|
Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai

Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|
Suchodus durobrivensis
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Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|
Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|
Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|
Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai

Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis
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Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|
Suchodus durobrivensis
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Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|
Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|
Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|
Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai

Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis
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Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|
Suchodus durobrivensis
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Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|
Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|
Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|
Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai

Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|
Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|
Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|
Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|
Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|Metriorhynchus' casamiquelai

Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis
-------
Image:Geosaurus BW.jpg|thumb|center|150px|Cricosaurus suevicus
Image:Dakosaurus BW.jpg|thumb|center|150px|Dakosaurus andiniensis
Image:Metriorhynchus superciliosum.jpg|thumb|center|150px|Thalattosuchus superciliosus
Image:Neptunidraco NT.jpg|thumb|center|150px|Neptunidraco ammoniticus
Image:Suchodus BW.jpg|thumb|center|150px|'Metriorhynchus' casamiquelai
Image:Suchodus durobrivense.jpg|thumb|center|150px|Suchodus durobrivensis