Tsylmosuchus
Tsylmosuchus is an extinct genus of proterosuchid archosauriform reptile known from Western Russia. Fossils referred to Tsylmosuchus occurred over a wide area in sediments corresponding to the Induan and Olenekian stages of the Early Triassic. Most of these fossils are fragmentary neck vertebrae which were originally reported as sharing similarities with crocodile-line archosaurs such as Mandasuchus. As a result, Tsylmosuchus was first described as part of the family Rauisuchidae, making it supposedly one of the oldest known archosaurs. However, its fragmentary remains do not show any of the distinguishing features of rauisuchids or even pseudosuchians in general, so Tsylmosuchus has more recently been interpreted as an indeterminate archosauriform. Although three species of Tsylmosuchus have been named, they lack diagnostic traits and are probably not distinct from each other. In 2023, Tsylmosuchus was reinterpreted as a proterosuchid, specifically a member of the subfamily Chasmatosuchinae.
Discovery
The genus is named after the Tsilma River, near where fossils have been found. Three species were named along with the original genus in 1990, their holotypes each comprising a single cervical vertebra and all of them only tentatively differentiated by their ages. The holotype of the type species T. jakovlevi was found near Ust'-Tsylma in the Komi Republic. This specimen and other referred material come from the Mezen' and Pechora river basins, which preserve sediments of the Sludkian and Ustmylian gorizonts. Some material once referred to the proterosuchid Chasmatosuchus magnus has since been referred to this species. T. donensis comes from the Donskaya Luka site in Volgograd Oblast, which preserves the Lipovskaya Formation of the Yarenskain Gorizont.Tsylmosuchus samariensis, described from a single site in Borskoi dating to the Rybinskian Gorizont, was synonymized with Chasmatosuchus rossicus by Ezcurra, who also made the Tsylmosuchus type species a nomen dubium. However, in 2023, Ezcurra reversed this decision and placed all three species in Tsylmosuchus again, and found the genus to be a proterosuchid.
Description
Tsylmosuchus is known primarily from vertebrae. These vertebrae have been compared to "rauisuchid" vertebrae, from the Manda Beds of Tanzania, which are now referred to Mandasuchus. Like several contemporary "rauisuchians", Tsylmosuchus has elongated cervical vertebrae. This would have given it a relatively long neck.Nesbitt suggested that elongated cervical vertebrae were present in numerous archosauriform clades, and that the length of the cervical vertebrae depended on the position in the presacral column, as seen in the poposauroid Arizonasaurus and the non-archosaurian archosauriform Guchengosuchus. The holotypes of each of the three species consist of a single cervical vertebra, which does not bear any other clear autapomorphies. Furthermore, the material referred to these species cannot be shown to belong to the same taxa as the holotypes. Thus, Nesbitt considered all three species to represent invalid archosauriforms.
Sennikov interpreted Tsylmosuchus as a member of the Ctenosauriscidae and also suggested that Scythosuchus may have been the same animal as Tsylmosuchus. In contrast, it was recovered as a member of the Proterosuchidae by Ezcurra, who reversed some of his previous reclassifications of the genus.