Salsoloideae
The Salsoloideae are a subfamily of the Amaranthaceae, formerly in family Chenopodiaceae.
Description
These are herbs, subshrubs, shrubs and some trees. Stems and leaves are often succulent. The ovary contains a spiral embryo. In most genera, scarious wings develop at the outside of the fruiting perianth, allowing for dispersal by the wind.In tribe Caroxyleae, the stamens have vesiculose anther appendages, discolor with anthers, that probably play a role for insect pollination. In tribe Salsoleae the anther appendages are absent or small and inconspicuous.
Distribution
The area with most species are the deserts and semideserts of Central-Asia and the Middle East. Distribution of the subfamily extends to the Mediterranean, to Middle-Europe, north and south Africa, and Australia, some species have also been introduced to America. Many species grow in dry habitats or tolerate salty soils, some are ruderals.Photosynthesis pathway
Salsoloideae are plants. Tribe Caroxyleae is exclusively of the NAD-malic enzyme subtype. Most Salsoleae also use the NADP-malic enzyme.Taxonomy
The type genus of subfamily Salsoloideae is Salsola L. According to recent research, the former classification did not reflect the phylogenetic relationship, so that the subfamily had to be reclassified; it is now split in two tribes, Caroxyleae and Salsoleae. The tribe Camphorosmeae is now treated in its own subfamily, Camphorosmoideae.Caroxyleae
Caroxyleae Akhani & E. H. Roalson Akhania, with 5 speciesCaroxylon Thunb., with 128 speciesClimacoptera Botsch., with 41 speciesHalanthium K.Koch, with 5 speciesHalarchon Bunge, with one species- * Halarchon vesiculosum BungeHalimocnemis C. A. Mey., with 27 species Halocharis Moq., with 7 speciesKaviria Akhani & E. H. Roalson, with 10 speciesNanophyton Less., with ca. 10 speciesOfaiston Raf., with one species
- * Ofaiston monandrum Moq.Petrosimonia Bunge, with 12 speciesPiptoptera Bunge, with one species
- * Piptoptera turkestana BungePhysandra Botsch., with one species
- * Physandra halimocnemis Botsch.Pyankovia Akhani & E. H. Roalson, with one species
- * Pyankovia brachiata Akhani & E. H. Roalson
Salsoleae
Anabasis L., with 29 speciesArthrophytum Schrenk, with 9 speciesCornulaca Delile, with 5 speciesCyathobasis Aellen, with one species:- * Cyathobasis fruticulosa AellenGirgensohnia Bunge ex Fenzl, with 4 speciesHalogeton C. A. Mey, with 5 species. Halothamnus Jaub. & Spach, with 21 speciesHaloxylon BungeHammada Iljin, with 12 speciesHoraninovia Fisch. & C. A. Mey, with 8 speciesLagenantha Chiov. Classification not sure. With 1-3 species.Noaea Moq., with 3 speciesNucularia Batt., Classification not sure. With one species:
- * Nucularia perrini Batt.Oreosalsola Rhaphidophyton Iljin, with one species
- * Rhaphidophyton regelii IljinSalsola L., with 56 species. Soda Sympegma BungeTraganum Del., with 2 speciesTraganopsis Maire et Wilczek, with one species
- * Traganopsis glomerata Maire & WilczekTurania Akhani & E. H. Roalson, with 4 speciesXylosalsola Tzvelev, with 4 species
Classification in subfamily not sure
Iljinia Korovin, with one species:- * ''Iljinia regelii''