Red-footed falcon


The red-footed falcon, formerly the western red-footed falcon, is a bird of prey. It belongs to the family Falconidae, the falcons. This bird is found in eastern Europe and Asia although its numbers are dwindling rapidly due to habitat loss and hunting. It is migratory, wintering in Africa. It is a regular wanderer to western Europe, and in August 2004 a red-footed falcon was found in North America for the first time on the island of Martha's Vineyard, Massachusetts.

Taxonomy

The Amur falcon was formerly included herein as a subspecies but it is nowadays considered distinct. Nonetheless, it is the present species' closest relative; their relationship to other falcons is more enigmatic. They appear morphologically somewhat intermediate between kestrels and hobbies and DNA sequence data has been unable to further resolve this question, mainly due to lack of comprehensive sampling. They might be closer to the merlin than to most other living falcons, or more generally related to this species and American falcons such as the American kestrel and the aplomado falcon.
The genus name Falco is from Late Latin falx, falcis, a sickle, referencing the claws of the bird. The species name vespertinus is Latin for "of evening" from vesper, "evening".

Description

It is a medium-small, long-winged species. The adult male is all blue-grey, except for his red undertail and legs; its underwings are uniformly grey. The female has a grey back and wings, orange head and underparts, and a white face with black eye stripe and moustaches.
Young birds are brown above and buff below with dark streaks, and a face pattern like the female. Red-footed falcons are in length with a wingspan of. The average mass is.

Distribution

The main areas of European distribution are in southern Russia and Ukraine. The species occurs in large numbers in Hungary, Romania and Serbia. Smaller populations exist in Italy, Bulgaria and Moldova, in Austria, Slovakia and in Belarus. The red-footed falcon also breeds irregularly in the Czech Republic and in the Baltic States, occasionally also in Germany. In Asia, the breeding occurrences run in a relatively narrow band somewhat south of the taiga eastwards to the upper Lena, where they touch the range of the sister species Falco amurensis east of the Baikal. The northern border lies between the 63° and 58° north latitude, the south border around 45° north latitude. It runs along the north slope of the Altai and then westward follows the transition of open pine forests into the treeless steppe areas of Central Asia. Further to the west, the breeding areas on the lower reaches of the Volga and on the north coast of the Caspian Sea reach Europe. The southernmost occurrences are in the steppe areas north of the Caucasus and in South Georgia. Some isolated breeding areas are located in northern Turkey. The red-footed falcon winters in southern and eastern Africa.

Breeding and behaviour

This falcon is a colonial breeder, reusing the old nests of corvids, such as rooks. It lays two to four eggs. Its maximum lifespan is 13.25 years in the wild and 18 years in captivity.

Azerbaijan

Red-footed falcons were witnessed copulating several times over a week-long period and "showed a strong preference for an old magpie nest" Frequent copulation was interspersed with joint inspections of the magpie nest. The morning after the aforementioned observations, the pair was again observed and they repeated their behaviour. Both then defended against an attack by a lesser kestrel. On this day neither bird spent much time in the nest, but always remained perched beside it. At the end of the week the researchers revisited the nest. The behaviour this time was that the female sat in the nest and the male perched outside but in the same tree. Further defensive responses to lesser kestrels were observed. The male red-footed falcon hunted for large insects and fed them to the female. Copulation then occurred. After copulation, the female returned to sit in the nest until the male fed her again. This behaviour continued for several days. This pair clearly showed territorial behaviour. The frequent copulations and lengthy stay of the female in the nest suggested incubation but the researchers did not check the nest.

Carpathian Basin

Researchers used a species distribution model for red-footed falcons. This model can play a crucial role in identifying key nesting sites for endangered species. According to the researchers, the red-footed falcon is officially listed as near-threatened, due to the drastic breeding population decline of the past decades. Red-footed falcons breed in colonies and in solitary pairs. They do not build a nest. In Hungary, the landscape scale distribution of rookeries remained stable, while the density and size of rookeries decreased and their location shifted to human settlements. Similar patterns were reported from other European countries. The reasons of rookery declines can be attributed to a large-scale persecution in the mid-80s resulting in a 90% population crash. Because of this, most potential breeding colonies for red-footed falcons disappeared, causing a decline in the number of breeding pairs. Conservation projects have established artificial nest box colonies to rectify this problem.
There is little evidence of this plan's effectiveness and also little evidence on recent population trends and distribution, from Northern Serbia, where 5–10% of the total European Union population is thought to breed. The researchers' main focus was on understanding the relationship between landscape scale habitat variables and red-footed falcon presence. The researchers found that the increase of natural grasslands had a positive effect on nest site presence while the increase of broad-leaved forests negatively influenced the probability of nest site presence. Their modelling approach proved successful in describing the landscape scale habitat composition of red-footed falcon breeding sites in the modelling area. Natural grasslands have a considerable impact on the probability of nest site presence. Therefore, locating their breeding sites or creating breeding sites by providing artificial colonies with the aid of our results has additional conservation and socio-economic benefits for wildlife and human population.

Serbia

Near the village of Melenci there were 22 red-footed falcon nests in 1991 in which offspring had been successfully brought up. The contents of four nests were checked daily from the start of incubation until the fledglings left the nests. Breeding success did not differ between the group of four nests disturbed by inspections and the remaining 18. Despite the fact that the ratio of parent bird presence and absence differed in the comparison of the four nests, red-footed falcon parents were usually found in the close surroundings, i.e. they attended their nest. The attendance of the birds to the nests differed between the sexes during incubation as well as hatching and the nestling period, which fact can be interpreted as a difference between the roles of females and males. In 59% of the cases it was the female, while in 41% it was the male bird that was sitting on the eggs. At the time of hatching it was mostly the female that was present in the nest, while after hatching it was only the female. Later on the adult birds usually took off from the branch supporting the nest, from neighbouring trees, or from abandoned rook nests, rather than from the nest itself. Females participated more times in the defence of the nest than males, yet the ratios of the studied types of behaviour did not differ between the sexes. However, significant difference appeared to be present between four nests when the distribution frequency of the three behaviour types was looked at. The distribution of the reaction types of the parent birds to disturbance was significantly different in the periods of incubation, hatching and nestling, respectively. As nesting proceeded and parental investment grew, the number of repellences and attacks increased proportionally.
The survey of population size and distribution of red-footed falcons proceeded in June–July 2000 and 2001 respectively, ten years after the first census. Data of only those nests were processed in which there was breeding. Breeding Success was calculated from the number of offspring per reproductive female. During the survey in Voivodina in 1990–1991 there were 308 and 124 pairs, respectively, whereas ten years later, in the year 2000 there were 116, and in 2001 only 61 pairs of red-footed falcons. Even if the marked fluctuations observed are not considered, the red-footed falcon population breeding in Voivodina shows a declining tendency. The south-western margin of the distribution area has moved towards the northeast by about 50–70 kilometres. More than 90% of the nesting sites, including the larger nesting colonies, are found in the Banat region, i.e. east of the Tisa River. More than 90% of the red-footed falcons continue to nest in rook colonies. No significant change has occurred in their breeding success.

Dietary habits

The red-footed falcon is a bird of prey with a diet consisting of a variety of insects, amphibians, reptiles, mammals and birds, such as great green bush-crickets, spadefoot toads, sand lizards, the common vole and bird nestlings, respectively. This bird's distinctive method of hunting is shared by the common kestrel. It regularly hovers, searching the ground below, then makes a short steep dive towards the target. When feeding their nestlings, the youngest nestlings receive the most food more frequently and more regularly. Chicks that are between 0 and 3 weeks old will get fed bigger prey like toads, lizards, bird nestlings, and great green bush-crickets, while nestlings over that age will get fed less frequently and with less variety. The diet of young nestlings consists mostly of Orthoptera and beetles with some vertebrates, while the diet of older nestlings mostly consists of Orthoptera and almost never of vertebrates.