Obrimini

The Obrimini are the most species-rich tribe of the Phasmatodea family of the Heteropterygidae native to Southeast Asia.

Description

The Obrimini differ from their sister tribe the Hoplocloniini by the structure of the secondary ovipositor at the abdomen of the females, which surrounds the actual ovipositor. It is formed in the Obrimini dorsally from the eleventh abdominal tergum called supraanal plate or epiproct, while in the Hoplocloniini it originated from the tenth tergum.

Distribution area

The distribution area of the Obrimini extends from Borneo to the east and includes the Philippines, Sulawesi, most of the Moluccas and New Guinea. Farthest east lies with Viti Levu, the main island of the Fiji group, the distribution area of Pterobrimus depressus.

Taxonomy

Brunner von Wattenwyl built in 1893 for the genera already described Obrimus, Hoploclonia, Tisamenus, Pylaemenes, Dares and Datames the tribe Obrimini. He placed these together with the genus Heteropteryx and the South American tribe Cladomorphini in the family Cladomorphidae. In the following years, basesd on this work, mostly genera of today's Obriminae and Dataminae were listed in this tribe, for example by [Josef Josef Redtenbacher (entomologist)|Redtenbacher (entomologist)|Josef Redtenbacher] in 1906. Lawrence Bruner raised the Obrimini to the rank of a family in 1915. James Abram Garfield Rehn and his son John William Holman Rehn divided the Obriminae, which they only referred to as a subfamily, into the tribes Obrimini and Datamini in 1939. Both tribes were transferred in 1953 by Klaus Günther to the subfamily Heteropteryginae. In 2004 Oliver Zompro raised this subfamily to the rank of family and the tribes contained in the rank of subfamilies or in the rank of a separate family. At the same time he built three tribes in the subfamily Obriminae. In addition to the Obrimini, these were the Miroceramiini and the Eubulidini. The latter was in 2016 by Frank H. Hennemann et al synonymized with the Obrimini. At the same time, the new tribe Tisamenini was established within the Obriminae. In a study based on genetic analysis, published in 2021, Sarah Bank et al. synonymized the tribe Miroceramiini and the tribe Tisamenini with the Obrimini and placed next to this the Hoplocloniini as the only other tribe in the subfamily Obriminae. In addition, a species previously referred to as Trachyaretaon sp. 'Negros' was identified as a representative of an as yet undescribed genus. Hennemann described this in 2023 alongside many other species of the tribe as Trachyaretaon negrosanon.

Genera with species

Sources:

Image	Genus	Species
	Aretaon Rehn, J.A.G. & Rehn, J.W.H., 1939	Aretaon asperrimus Aretaon muscosus
	Armadolides Hennemann, 2023	Armadolides manobo
	Brasidas Rehn, J.A.G. & Rehn, J.W.H., 1939	Brasidas bakeri Brasidas cavernosus Brasidas foveolatus Brasidas lacerta Brasidas malaki Hennemann, 2023Brasidas manobo Hennemann, 2023Brasidas rehni Hennemann, 2023Brasidas samarensis Rehn, J.A.G. & Rehn, J.W.H., 1939Brasidas viscayanus Rehn, J.A.G. & Rehn, J.W.H., 1939Brasidas waray Hennemann, 2023
	Eubulides Stål, 1877	Eubulides alutaceus Stål, 1877Eubulides blaan Hennemann, 2023Eubulides constanti Hennemann, 2023Eubulides igorrote Rehn, J.A.G. & Rehn, J.W.H., 1939Eubulides lumawigi Hennemann, 2023Eubulides taylori Rehn, J.A.G. & Rehn, J.W.H., 1939Eubulides timog Hennemann, 2023
	Heterocopus Redtenbacher, 1906	Heterocopus leprosus Redtenbacher, 1906
	Mearnsiana Rehn, J.A.G. & Rehn, J.W.H., 1939	Mearnsiana bullosa Rehn, J.A.G. & Rehn, J.W.H., 1939Mearnsiana maranao Hennemann, 2023
	Miroceramia Günther, 1934	Miroceramia westwoodii
	Obrimus Stål, 1875	Obrimus bicolanus Rehn, J.A.G. & Rehn, J.W.H., 1939Obrimus bufo Obrimus mesoplatus Obrimus uichancoi Rehn, J.A.G. & Rehn, J.W.H., 1939
	Pterobrimus Redtenbacher, 1906	Pterobrimus depressus Redtenbacher, 1906
	Stenobrimus Redtenbacher, 1906	Stenobrimus bolivari Redtenbacher, 1906Stenobrimus lumad Lit & Eusebio, 2010Stenobrimus pilipinus Eusebio, Lit & Lucañas, 2023Stenobrimus tagalog Rehn, J.A.G. & Rehn, J.W.H., 1939
	Sungaya Zompro, 1996	Sungaya aeta Hennemann, 2023Sungaya dumagat Hennemann, 2023Sungaya ibaloi Hennemann, 2023Sungaya inexpectata Zompro, 1996
	Theramenes Stål, 1875	Theramenes dromedarius Stål, 1877Theramenes exiguus Hennemann & Conle, 2003Theramenes letiranti Hennemann, 2023Theramenes mandirigma Zompro & Eusebio, 2001Theramenes olivaceus
	Tisamenus Stål, 1875	Tisamenus alviolanus Lit & Eusebio, 2010Tisamenus armadillo Redtenbacher, 1906Tisamenus asper Tisamenus cervicornis Ignacio Bolívar, 1890Tisamenus charestae Hennemann & Le Tirant, 2025Tisamenus clotho Tisamenus deplanatus Tisamenus draconina Tisamenus fratercula Tisamenus hebardi Tisamenus heitzmanni Hennemann, 2025Tisamenus hystrix Tisamenus irenoliti Hennemann, 2025Tisamenus kalahani Lit & Eusebio, 2005Tisamenus lachesis Tisamenus makinis Hennemann, 2025Tisamenus malawak Hennemann, 2025Tisamenus napalaki Hennemann, 2025Tisamenus polillo Tisamenus ranarius Tisamenus serratorius Stål, 1875Tisamenus spadix Tisamenus summaleonilae Lit & Eusebio, 2005Tisamenus tagalog Tisamenus trapezoides Hennemann, 2025
	Trachyaretaon Rehn, J.A.G. & Rehn, J.W.H., 1939	Trachyaretaon bresseeli Hennemann, 2023Trachyaretaon carmelae Lit & Eusebio, 2005Trachyaretaon echinatus Trachyaretaon gatla Zompro, 2004Trachyaretaon maliit Hennemann, 2023Trachyaretaon mangyan Hennemann, 2023Trachyaretaon nakatago Hennemann, 2023Trachyaretaon negrosanon Hennemann, 2023Trachyaretaon tumandok Hennemann, 2023