Llanos


The Llanos is a vast tropical grassland plain situated to the east of the Andes in Colombia and Venezuela, in northwestern South America. It is an ecoregion of the tropical and subtropical grasslands, savannas, and shrublands biome.

Geography

The Llanos occupy a lowland that extends mostly east and west. The Llanos are bounded on the west and northwest by the Andes, and on the north by the Venezuelan Coastal Range. The Guiana Highlands are to the southeast, and the Negro-Branco moist forests are to the southwest. To the east the Orinoco wetlands and Orinoco Delta swamp forests occupy the Orinoco Delta. The Llanos' main river is the Orinoco, which runs from west to east through the ecoregion and forms part of the Colombia–Venezuela border. The Orinoco is the major river system of Venezuela.
A 2017 assessment found that 105,323 km², or 28%, of the ecoregion, is in protected areas. Protected areas include Aguaro-Guariquito National Park, Cinaruco-Capanaparo National Park, Tortuga Arrau Reserve, and Caño Guaritico Wildlife Refuge in Venezuela, and El Tuparro National Natural Park in Colombia.

Climate

The ecoregion has a tropical savanna climate that grades into a tropical monsoon climate in the Colombian Llanos. Rainfall is seasonal, with a rainy season from April to November, and a dry season between December and March. The wettest months are typically June and July. Rainfall varies across the ecoregion, from up to per year in the southwest, in Apure State, and per year in Monagas State in the northeast. Mean annual temperature is, and the average monthly temperature varies little throughout the year; the lowest-temperature months are only cooler than the hottest months.

Flora

The plant communities in the Llanos include open grasslands, savanna with scattered trees or clumps of trees, and small areas of forest, typically gallery forests along rivers and streams. There are seasonally flooded grasslands and savannas and grasslands and savannas that remain dry throughout the year.
The llano alto grasslands and savannas are characterized by grasses and shrubs 30–100 cm high, forming tussocks 10 to 30 cm apart. Soils are typically sandy and nutrient-poor. Llano alto covers approximately two-thirds of the Venezuelan llanos and is also widespread in the Colombian llanos. Grasses of genus Trachypogon are predominant, and species include Trachypogon plumosus, T. vestitus, Axonopus canescens, A. anceps, Andropogon selloanus, Aristida spp., Leptocoryphium lanatum, Paspalum carinatum, Sporobolus indicus, and S. cubensis, and sedges in the genera Rhynchospora and Bulbostylis. Shrubs and herbs are most commonly legumes in the genera Mimosa, Cassia, Desmodium, Eriosema, Galactia, Indigofera, Phaseolus, Stylosanthes, Tephrosia, and Zornia. The trees manteco, chaparro, and alcornoque are the most common, growing either as scattered trees or in woodland patches known as matas which range in area from 12 meters in diameter up to a hectare.
During the rainy season, parts of the Llanos can flood up to a meter. This turns some savannas and grasslands into temporary wetlands, comparable to the Pantanal of central South America. This flooding creates habitat for water birds and other wildlife. These seasonally flooded grasslands and savannas, known as llano bajo, typically have richer soils. They are characterized by the grass Paspalum fasciculatum. Trees include the palm Copernicia tectorum and gallery forest species.
Gallery forests include evergreen seasonally flooded forests, and semi-deciduous forests on higher ground. Morichales are seasonally flooded forests characterized by the moriche palm. Vegas are seasonally flooded evergreen forests found along the Orinoco and its tributaries. Trees form a canopy 8 to 20 meters high and include Inga spp., Combretum frangulifolium, Gustavia augusta, Pterocarpus sp., Pterocarpus dubius, Spondias mombin, and Copaifera pubiflora. Semi-deciduous forests occur above flood level and form a canopy 12 to 15 meters high. Common trees include Tabebuia billbergii, Godmania aesculifolia, Cassia moschata, Spondias mombin, Copaifera pubiflora, Bourreria cumanensis, Cordia spp., Bursera simaruba, Cochlospermum vitifolium, Hura crepitans, and Acacia glomerosa.
"Matorrales" are deciduous and semi-deciduous shrublands 5 to 8 meters high which cover large areas in the central Venezuelan llanos and may be a form of secondary vegetation in areas that were formerly dry deciduous forest. Typical shrubs are Bourreria cumanensis, Randia aculeata, Godmania aesculifolia, Pereskia guamacho, Prosopis spp., Xylosma benthamii, Erytroxylum sp., and Cereus hexagonus.

Fauna

Mammals of the grassland and savanna include white-tailed deer, giant anteater, capybara, giant armadillo, Eastern cottontail, Alston's cotton rat, Hispid cotton rat, Zygodontomys brevicauda, and Oecomys bicolor.
The gallery forests are home to more diverse large and medium-sized mammals, including collared peccary, white-lipped peccary, South American tapir, white-tailed deer, red brocket, wedge-capped capuchin, Venezuelan red howler, large rodents like the lowland paca, agoutis, and Brazilian porcupine, and large cats like the puma, jaguar, and ocelot. The endangered giant otter lives along the Orinoco and its tributaries. Some of the largest jaguars in the world are found in the Llanos, with average weights of over for males. The Llanos long-nosed armadillo and the short-tailed opossum Monodelphis orinoci are endemic to the Llanos.
The wetlands support around 70 species of water birds, including the scarlet ibis. A large portion of the distribution of the sharp-tailed ibis and white-bearded flycatcher is in the Llanos. Native reptiles include the Orinoco crocodile, spectacled caiman, Green anaconda, and Arrau turtle.

Indigenous peoples

Indigenous peoples of the Llanos include the Guahibo in the western Llanos of Colombia and Venezuela, and the Yaruro in the eastern Llanos in Venezuela.

Economy

The primary economic activity since the Spanish colonial era is the herding of millions of cattle. An 1856 watercolor by Manuel María Paz depicts sparsely populated open grazing lands with cattle and palm trees. The term llanero became synonymous with the cowhands that took care of the herds and had some cultural similarities with the gauchos of the Pampas or the vaqueros of Spanish and Mexican Texas. Decades of extensive cattle raising has altered the ecology of the Llanos. Grasslands and savannas are frequently burned to make them more suitable for grazing and eliminate trees and shrubs. Non-native grasses have been introduced for cattle fodder, including the African grass Melinis minutiflora, and now cover large areas. Agriculture, particularly rice and maize, now cover extensive areas, including rice fields in former seasonal wetlands.
The governments of Venezuela and Colombia have developed a strong oil and gas industry in Arauca, Casanare, Guárico, Anzoátegui, Apure, and Monagas. The Orinoco Belt, entirely in Venezuelan territory, consists of large deposits of extra heavy crude. The Orinoco Belt oil sands are known to be one of the largest, behind that of the Athabasca Oil Sands in Alberta, Canada. Venezuela's non-conventional oil deposits of about, found primarily in the Orinoco oil sands, are estimated to approximately equal the world's conventional oil reserves.

Cities

Colombia

  • Acacías
  • Arauca, Arauca
  • Gaviotas
  • Maní, Casanare
  • Orocué
  • Paz de Ariporo
  • Puerto Carreño
  • Inírida
  • Puerto López, Meta
  • San José del Guaviare
  • Saravena
  • Tame
  • Villavicencio
  • Yopal
  • Fortul

    Venezuela

  • Acarigua
  • Araure
  • Barinas
  • Calabozo
  • Caripito
  • El Tigre
  • Guanare
  • Maturín
  • Puerto Ayacucho
  • Sabaneta
  • San Carlos
  • San Fernando de Apure
  • Tucupita
  • Valle de la Pascua