Hybridization in pines

Both naturally and artificially occurring pine species (Pinus) can hybridize, combining their genetic material and sometimes creating hybrids that can be more or less vigorous than their parent species. An example of a naturally occurring hybrid pine is Pinus × sondereggeri, a naturally occurring cross between loblolly pine and longleaf pine. An example of the many artificial hybrids is Pinus lambertiana × ''P. armandii.''

Subgenus ''Pinus''

Includes both natural and artificial pine hybrids.

Subsection ''Pinus''

Pinus nigra × P. resinosa – Austrian pine × red pine
Pinus densiflora × P. nigra ''– Japanese Red pine × Austrian pine
Pinus × densithunbergii, Pinus densiflora × P. thunbergii –'' Japanese Red pine × Japanese black pine
Pinus × neilreichiana, Pinus nigra × P. sylvestris – Neilreich pine
Pinus × rhaetica, Pinus mugo × P. sylvestris – Rhætic pine

Subsection ''Ponderosae''

Pinus ponderosa var. ponderosa × P. arizonica – Ponderosa pine × Arizona pine
Pinus ponderosa var. ponderosa × P. montezumae – Ponderosa pine × Montezuma pine
Pinus ponderosa var. scopulorum × P. montezumae – Rocky Mountain ponderosa pine × Montezuma pine
Pinus engelmannii × P. montezumae – Apache pine × Montezuma pine
Pinus jeffreyi × P. montezumae – Jeffrey pine × Montezuma pine
Pinus ponderosa var. ponderosa × P. engelmannii – Ponderosa pine × Apache pine
Pinus engelmannii × P. arizonica – Apache pine × Arizona pine
Pinus engelmannii × P. ponderosa var. scopulorum – Apache pine × Rocky Mountain ponderosa pine
Pinus jeffreyi × P. ponderosa var. ponderosa – Jeffrey pine × ponderosa pine
Pinus jeffreyi × P. ponderosa var. scopulorum – Jeffrey pine × Rocky Mountain ponderosa pine
Pinus jeffreyi × P. ponderosa var. washoensis – Jeffrey pine × Washoe pine
Pinus. ponderosa var. washoensis × P. ponderosa var. ponderosa – Washoe pine × ponderosa pine
Pinus. ponderosa var. washoensis × P. ponderosa var. scopulorum – Washoe pine × Rocky Mountain ponderosa pine
Pinus jeffreyi × P. coulteri – Jeffrey pine × Coulter pine

Subsection ''Contortae''

Pinus × murraybanksiana, Pinus contorta var. murrayana × P. banksiana – Murraybanks pine
Pinus contorta var latifolia × P. banksiana – Rocky Mountain Lodgepole Pine × Jack Pine
Pinus virginiana × P. clausa – Virginia pine × Sand pine

Subsection ''Australes''

Pinus × sondereggeri, Pinus palustris × P. taeda – Sonderegger's pine
Pinus elliottii × P. palustris – Slash pine × longleaf pine
Pinus elliottii × P. taeda – Slash pine × loblolly pine
Pinus echinata × P. elliottii – Shortleaf pine × slash pine
Pinus echinata × P. taeda – Shortleaf pine × loblolly pine
Pinus pungens × P. echinata – Table Mountain pine × shortleaf pine
Pinus rigida × P. echinata – Pitch pine × shortleaf pine
Pinus x rigitaeda, Pinus rigida × P. taeda – Pitlolly pine
Pinus rigida × P. serotina – Pitch pine × pond pine
Pinus patula × P. greggii – Mexican weeping pine × Gregg pine
Pinus patula × P. radiata – Mexican weeping pine × Monterey pine
Pinus attenuata × P. radiata – KMX pine
Pinus attenuata × P. muricata – Knobcone pine × bishop pine

Subgenus ''Strobus''

Includes both natural and artificial pine hybrids. There have been external reports of pine hybrids in the subgenus Strobus.

Subsection ''Cembroides''

Pinus quadrifolia × P. monophylla subsp. californiarum ''– Parry pinyon × California single-leaf pinyon
Pinus quadrifolia × P. monophylla – Parry pinyon × single-leaf pinyon
Pinus monophylla × P. edulis'' – Single-leaf pinyon × Colorado pinyon

Subsection ''Strobus''

Pinus pumila × P. sibirica – Japanese stone pine × Siberian pine
Pinus × hakkodensis, Pinus parviflora var. pentaphylla × P. pumila – Hakkoda pine
Pinus × holfordiana, Pinus ayacahuite × P. wallichiana – Holford pine Pinus × hunnewellii, Pinus parviflora × P. strobus – Hunnewell's white pine Pinus monticola × P. parviflora – Western white pine × Japanese white pine
Pinus lambertiana × P. armandii – Sugar pine × Armand pine
Pinus lambertiana × P. koraiensis – Sugar pine × Korean pine
Pinus monticola × P. strobiformis – Western white pine × southwestern white pine
Pinus monticola × P. flexilis – Western white pine × limber pine
Pinus monticola × P. strobus – Western white pine × eastern white pine
Pinus monticola × P. peuce – Western white pine × Balkan pine
Pinus peuce × P. strobus – Balkan pine × eastern white pine
Pinus peuce × P. parviflora – Balkan pine × Japanese white pine
Pinus flexilis × P. wallichiana – Limber pine × Himalayan pine
Pinus flexilis × P. strobus – Limber pine × eastern white pine
Pinus flexilis × P. ayacahuite – Limber pine × Mexican white pine
Pinus ayacahuite × P. strobus – Mexican white pine × eastern white pine
Pinus × schwerinii, Pinus strobus × P. wallichiana – Schwerin's white pine
Pinus monticola × P. wallichiana – Western white pine × Himalayan pine
Pinus cembra × P. sibirica – Swiss stone pine × Siberian pine
Pinus cembra × P. albicaulis – Swiss stone pine × whitebark pine
Pinus albicaulis × P. flexilis – Whitebark pine × limber pine
Pinus sibirica × P. cembra – Siberian pine × Swiss stone pine
Pinus sibirica × P. koraiensis – Siberian pine × Korean pine
Pinus armandii × P. koraiensis – Armand pine × Korean pine
Pinus strobus × P. ayacahuite – Eastern white pine × Mexican white pine