Etheostoma
Etheostoma is a genus of small freshwater fish in the family Percidae, and within the sub-family Etheostomatinae, native to North America. Most are restricted to the United States, but species are also found in Canada and Mexico. They are commonly known as darters, although the term "darter" is shared by several other genera. Many can produce alarm pheromones that serve to warn nearby fish in case of an attack.
Species
The 146 recognized species in this genus are arranged into 18 subgenera. Two former subgenera, Allohistium and Nothonotus have been promoted to full genera.- Subgenus Belophlox Fowler, 1947
- * Etheostoma fricksium Hildebrand, 1923
- * Etheostoma mariae
- * Etheostoma okaloosae
- Sugbenus Boleosoma DeKay, 1842
- * Etheostoma longimanum D. S. Jordan, 1888
- * Etheostoma maculaticeps
- * Etheostoma nigrum Rafinesque, 1820
- * Etheostoma olmstedi D. H. Storer, 1842
- * Etheostoma perlongum C. L. Hubbs & Raney, 1946)
- * Etheostoma podostemone D. S. Jordan & O. P. Jenkins, 1889
- * Etheostoma susanae
- * Etheostoma vexillare
- Sugbenus Catonotus Agassiz, 1854
- * Etheostoma barbouri Kuehne & J. W. Small, 1971
- * Etheostoma basilare Page, Hardman & Near, 2003
- * Etheostoma brevispinum
- * Etheostoma chienense Page & Ceas, 1992
- * Etheostoma corona Page & Ceas, 1992
- * Etheostoma crossopterum Braasch & Mayden, 1985
- * Etheostoma cumberlandicum D. S. Jordan & Swain, 1883
- * Etheostoma derivativum Page, Hardman & Near, 2003
- * Etheostoma flabellare Rafinesque, 1819
- * Etheostoma forbesi Page & Ceas, 1992
- * Etheostoma kennicotti
- * Etheostoma lemniscatum Blanton, 2008
- * Etheostoma marmorpinnum Blanton & R. E. Jenkins, 2008
- * Etheostoma nebra Near & M. R. Thomas, 2015
- * Etheostoma neopterum W. M. Howell & Dingerkus, 1978
- * Etheostoma nigripinne Braasch & Mayden, 1985
- * Etheostoma obeyense Kirsch, 1892
- * Etheostoma olivaceum Braasch & Page, 1979
- * Etheostoma oophylax Ceas & Page, 1992
- * Etheostoma percnurum R. E. Jenkins, 1994
- * Etheostoma pseudovulatum Page & Ceas, 1992
- * Etheostoma sitikuense Blanton, 2008
- * Etheostoma smithi Page & Braasch, 1976
- * Etheostoma squamiceps D. S. Jordan, 1877
- * Etheostoma striatulum Page & Braasch, 1977
- * Etheostoma virgatum
- * Etheostoma xanthovum Wood & Near, 2023
- Subgenus Doration D.S. Jordan, 1929
- * Etheostoma akatulo Layman & Mayden, 2009
- * Etheostoma clinton Mayden & Layman, 2012
- * Etheostoma gore Mayden & Layman, 2012
- * Etheostoma jessiae
- * Etheostoma jimmycarter Mayden & Layman, 2012
- * Etheostoma meadiae
- * Etheostoma obama Mayden & Layman, 2012
- * Etheostoma stigmaeum
- * Etheostoma teddyroosevelt Mayden & Layman, 2012
- Subgenus Etheostoma
- * Etheostoma blennioides Rafinesque, 1819
- * Etheostoma blennius C. H. Gilbert & Swain, 1887
- * Etheostoma gutselli
- * Etheostoma histrio D. S. Jordan & C. H. Gilbert, 1887
- * Etheostoma inscriptum
- * Etheostoma lynceum O. P. Hay, 1885
- * Etheostoma rupestre C. H. Gilbert & Swain, 1887
- * Etheostoma swannanoa D. S. Jordan & Evermann, 1889
- * Etheostoma sequatchiense Burr, 1979
- * Etheostoma thalassinum
- * Etheostoma zonale
- Subgenus Fuscatelum Page, 1981
- * Etheostoma parvipinne C. H. Gilbert & Swain, 1887
- * Etheostoma phytophilum Henry L. [Bart Jr.|Bart] & M. S. Taylor, 1999
- Subgenus Hololepis Frederic [Ward Putnam|Putnam], 1863
- * Etheostoma collis
- * Etheostoma fusiforme
- *Etheostoma gracile
- * Etheostoma serrifer
- * Etheostoma zonifer C. L. Hubbs & Cannon, 1935)
- Subgenus Ioa D.S. Jordan & Brayton, 1878
- * Etheostoma vitreum
- Subgenus Litocara R.M. Bailey, 1948
- * Etheostoma nianguae C. H. Gilbert & Meek, 1887
- * Etheostoma sagitta
- * Etheostoma spilotum C. H. Gilbert, 1887
- Subgenus Microperca Putnam, 1863
- * Etheostoma fonticola
- * Etheostoma microperca D. S. Jordan & C. H. Gilbert, 1888
- * Etheostoma proeliare
- Subgenus Mooreichthys R.R. Stephens, K.A. Johnson & M.P. Grady, 2014
- * Etheostoma sellare
- Sugenus Oligocephalus Girard, 1859
- * Etheostoma artesiae
- * Etheostoma asprigene
- * Etheostoma australe D. S. Jordan, 1889
- * Etheostoma bison Ceas & Page, 1997
- * Etheostoma burri Ceas & Page, 1997
- * Etheostoma caeruleum D. H. Storer, 1845
- * Etheostoma collettei Birdsong & L. W. Knapp, 1969
- * Etheostoma cyanorum
- * Etheostoma ditrema Ramsey & Suttkus, 1965
- * Etheostoma exile
- * Etheostoma fragi Distler, 1968
- * Etheostoma grahami
- * Etheostoma hopkinsi
- * Etheostoma kantuckeense Ceas & Page, 1997
- * Etheostoma lawrencei Ceas & Burr, 2002
- * Etheostoma lepidum
- * Etheostoma lugoi S. M. Norris & W. L. Minckley, 1997
- * Etheostoma luteovinctum C. H. Gilbert & Swain, 1887
- * Etheostoma nuchale W. M. Howell & R. D. Caldwell, 1965
- * Etheostoma pottsii
- * Etheostoma radiosum
- * Etheostoma segrex S. M. Norris & W. L. Minckley, 1997
- * Etheostoma spectabile
- * Etheostoma swaini
- * Etheostoma tecumsehi Ceas & Page, 1997
- * Etheostoma thompsoni Suttkus, Bart & Etnier, 2012
- * Etheostoma uniporum Distler, 1968
- * Etheostoma whipplei
- Subgenus Ozarka J.D. Williams & Robison, 1981
- * Etheostoma autumnale Mayden, 2010
- * Etheostoma boschungi Wall & J. D. Williams, 1974
- * Etheostoma cragini C. H. Gilbert, 1885
- * Etheostoma mihileze Mayden, 2010
- * Etheostoma pallididorsum Distler & Metcalf, 1962
- * Etheostoma punctulatum
- * Etheostoma trisella R. M. Bailey & W. J. Richards, 1963
- Subgenus Poecilichthys Agassiz, 1854
- * Etheostoma erythrozonum Switzer & R. M. Wood, 2009
- * Etheostoma euzonum
- * Etheostoma kanawhae
- * Etheostoma osburni
- * Etheostoma tetrazonum
- * Etheostoma variatum Kirtland, 1838
- Subgenus Psychromaster D.S. Jordan & Evermann, 1896
- * Etheostoma tuscumbia C. H. Gilbert & Swain, 1887
- Subgenus Ulocentra D.S. Jordan, 1878 “Snubnose darters”
- * Etheostoma atripinne
- * Etheostoma baileyi Page & Burr, 1982
- * Etheostoma barrenense Burr & Page, 1982
- * Etheostoma bellator Suttkus & R. M. Bailey, 1993
- * Etheostoma birminghamense Brownstein, Kim, Wood, Alley, Stokes & Near, 2025
- * Etheostoma brevirostrum Suttkus & Etnier, 1991
- * Etheostoma cervus Powers & Mayden, 2003
- * Etheostoma chermocki Boschung, Mayden & Tomelleri, 1992
- * Etheostoma colorosum Suttkus & R. M. Bailey, 1993
- * Etheostoma coosae
- * Etheostoma cyanoprosopum Near & Kozal, 2017
- * Etheostoma duryi Henshall, 1889
- * Etheostoma etnieri Bouchard, 1977
- * Etheostoma faulkneri Sterling & Warren, 2020
- * Etheostoma flavum Etnier & R. M. Bailey, 1989
- * Etheostoma gurleyense Brownstein, Kim, Wood, Alley, Stokes & Near, 2025
- * Etheostoma kimberlae Mayden, 2025
- * Etheostoma lachneri Suttkus & R. M. Bailey, 1994
- * Etheostoma michellae Mayden, 2025
- * Etheostoma orientale Powers & Mayden, 2007
- * Etheostoma planasaxatile Powers & Mayden, 2007
- * Etheostoma pyrrhogaster R. M. Bailey & Etnier, 1988
- * Etheostoma rafinesquei Burr & Page, 1982
- * Etheostoma ramseyi Suttkus & R. M. Bailey, 1994
- * Etheostoma raneyi Suttkus & Bart, 1994
- * Etheostoma scotti Bauer, Etnier & Burkhead, 1995
- * Etheostoma simoterum
- * Etheostoma tallapoosae Suttkus & Etnier, 1991
- * Etheostoma zonistium R. M. Bailey & Etnier, 1988
- Subgenus Vaillantia D.S. Jordan, 1878
- * Etheostoma chlorosoma
- * Etheostoma davisoni O. P. Hay, 1885
- Subgenus Villora Hubbs & Cannon, 1935
- * Etheostoma edwini