Dog
The dog is a domesticated descendant of wolves. Also called the domestic dog, it was selectively bred during the Late Pleistocene by hunter-gatherers. Dogs and the modern gray wolf share a common ancestor. Dogs were the first species to be domesticated over 14,000 years ago, before the development of agriculture, though genetic studies suggest the domestication process may have begun over 25,000 years ago. Due to their long association with humans, dogs have gained the ability to thrive on a starch-rich diet that would be inadequate for other canids.
Dogs have been bred for desired behaviors, sensory capabilities, and physical attributes. Dog breeds vary widely in shape, size, and color. They have the same number of bones, powerful jaws that house around 42 teeth, and well-developed senses of smell, hearing, and sight. Compared to humans, dogs possess a superior sense of smell and hearing, but inferior visual acuity. Dogs perform many roles for humans, such as hunting, herding, pulling loads, protection, companionship, therapy, aiding disabled people, and assisting police and the military.
Communication in dogs includes eye gaze, facial expression, vocalization, body posture, and chemical communication. They mark their territories by urinating on them, which is more likely when entering a new environment. Over the millennia, dogs have uniquely adapted to human behavior; this adaptation includes being able to understand and communicate with humans. As such, the human–canine bond has been a topic of frequent study, and dogs' influence on human society has given them the sobriquet of "man's best friend".
The global dog population is estimated at 700 million to 1 billion, distributed around the world. The dog is the most popular pet in the United States, present in 34–40% of households. Developed countries make up approximately 20% of the global dog population, while around 75% of dogs are estimated to be from developing countries, mainly in the form of feral and street dogs.
Taxonomy
Dogs are domesticated members of the family Canidae. They are classified as a subspecies of Canis lupus, along with wolves and dingoes. Genetic studies show that dogs likely diverged from wolves between 27,000 and 40,000 years ago. Dogs were domesticated from wolves at least 14,000 years ago by hunter-gatherers, before the development of agriculture; the remains of the Bonn–Oberkassel dog, buried alongside humans between 14,000 and 15,000 years ago, are the earliest to be conclusively identified as a domesticated dog. The dingo and the related New Guinea singing dog resulted from the geographic isolation and feralization of dogs in Oceania over 8,000 years ago.Dogs, wolves, and dingoes have sometimes been classified as separate species. In 1758, the Swedish botanist and zoologist Carl Linnaeus assigned the genus name Canis to the domestic dog, the wolf, and the golden jackal in his book, Systema Naturae. He classified the domestic dog as Canis familiaris and, on the next page, classified the grey wolf as Canis lupus. Linnaeus considered the dog to be a separate species from the wolf because of its upturning tail, which is not found in any other canid. In the 2005 edition of Mammal Species of the World, mammalogist W. Christopher Wozencraft listed the wolf as a wild subspecies of Canis lupus and proposed two additional subspecies: familiaris, as named by Linnaeus in 1758, and dingo, named by Meyer in 1793. Wozencraft included hallstromi as another name for the dingo. This classification was informed by a 1999 mitochondrial DNA study.
The classification of dingoes is disputed and a political issue in Australia. Classifying dingoes as wild dogs simplifies reducing or controlling dingo populations that threaten livestock. Treating dingoes as a separate species allows conservation programs to protect the dingo population. Dingo classification affects wildlife management policies, legislation, and societal attitudes. In 2019, a workshop hosted by the IUCN/Species Survival Commission's Canid Specialist Group considered the dingo and the New Guinea singing dog to be feral Canis familiaris. Therefore, it did not assess them for the IUCN Red List of threatened species.
Domestication
The earliest remains generally accepted to be those of a domesticated dog were discovered in Bonn-Oberkassel, Germany. Contextual, isotopic, genetic, and morphological evidence shows that this dog was not a local wolf. Dated to 14,223 years ago, the dog was found buried along with a man and a woman; all three had been sprayed with red hematite powder and buried under large, thick basalt blocks. The dog had survived canine distemper as a puppy - survival without intensive human care is unlikely, and occurred at an age before the dog could have had any utilitarian use. This timing indicates that the dog was the first species to be domesticated during the hunter-gatherer era, predating agriculture. Earlier remains dating back 30,000 years have been described as Paleolithic dogs, but their status as dogs or wolves remains debated because considerable morphological diversity existed among wolves during the Late Pleistocene.DNA sequences show that all ancient and modern dogs share a common ancestry and descended from an ancient, extinct wolf population that was distinct from any modern wolf lineage. Some studies have posited that all living wolves are more closely related to each other than to dogs, while others have suggested that dogs are more closely related to modern Eurasian wolves than to American wolves.
The dog is a domestic animal that likely travelled a commensal pathway into domestication. The questions of when and where dogs were first domesticated remains uncertain. Genetic studies suggest a domestication process commencing over 25,000 years ago, in one or several wolf populations in either Europe, the high Arctic, or eastern Asia. In 2021, a literature review of the current evidence infers that the dog was domesticated in Siberia 23,000 years ago by ancient North Siberians, then later dispersed eastward into the Americas and westward across Eurasia, with dogs likely accompanying the first humans to inhabit the Americas. Some studies have suggested that the extinct Japanese wolf is closely related to the ancestor of domestic dogs.
In 2018, a study identified 429 genes that differed between modern dogs and modern wolves. As the differences in these genes could also be found in ancient dog fossils, these were regarded as being the result of the initial domestication and not from recent breed formation. These genes are linked to neural crest and central nervous system development. These genes affect embryogenesis and can confer tameness, smaller jaws, floppy ears, and diminished craniofacial development, which distinguish domesticated dogs from wolves and are considered to reflect domestication syndrome. The study concluded that during early dog domestication, the initial selection was for behavior. This trait is influenced by those genes which act in the neural crest, which led to the phenotypes observed in modern dogs.
Breeds
There are around 450 official dog breeds, the most of any mammal. Dogs began diversifying in the Victorian era, when humans took control of their natural selection. Most breeds were derived from small numbers of founders within the last 200 years. Since then, dogs have undergone rapid phenotypic change and have been subjected to artificial selection by humans. The skull, body, and limb proportions between breeds display more phenotypic diversity than can be found within the entire order of carnivores. These breeds possess distinct traits related to morphology, which include body size, skull shape, tail phenotype, fur type, and colour. As such, humans have long used dogs for their desirable traits to complete or fulfill a certain work or role. Their behavioural traits include guarding, herding, hunting, retrieving, and scent detection. Their personality traits include hypersocial behavior, boldness, and aggression. Present-day dogs are dispersed around the world. An example of this dispersal is the numerous modern breeds of European lineage during the Victorian era.Anatomy and physiology
Size and skeleton
Dogs are extremely variable in size, ranging from one of the largest breeds, the Great Dane, at and, to one of the smallest, the Chihuahua, at and. All healthy dogs, regardless of their size and type, have the same number of bones, although there is significant skeletal variation between dogs of different types. The dog's skeleton is well adapted for running; the vertebrae on the neck and back have extensions for back muscles, consisting of epaxial muscles and hypaxial muscles, to connect to; the long ribs provide room for the heart and lungs; and the shoulders are unattached to the skeleton, allowing for flexibility.Compared to the dog's wolf-like ancestors, selective breeding since domestication has seen the dog's skeleton increase in size for larger types such as mastiffs and miniaturised for smaller types such as terriers; dwarfism has been selectively bred for some types where short legs are preferred, such as dachshunds and corgis. Most dogs naturally have 26 vertebrae in their tails, but some with naturally short tails have as few as three.
The dog's skull has identical components regardless of breed type, but there is significant divergence in terms of skull shape between types. The three basic skull shapes are the elongated dolichocephalic type as seen in sighthounds, the intermediate mesocephalic or mesaticephalic type, and the very short and broad brachycephalic type exemplified by mastiff type skulls. The jaw contains around 42 teeth, and it has evolved for the consumption of flesh. Dogs use their carnassial teeth to cut food into bite-sized chunks, more especially meat.