Colydiinae


Colydiinae is a subfamily of beetles, commonly known as cylindrical bark beetles. They have been treated historically as a family Colydiidae, but have been moved into the Zopheridae, where they constitute the bulk of the diversity of the newly expanded family, with about 140 genera worldwide. They are diverse for example in the Australian region, from where about 35 genera are known; in Europe, though, only 20 genera are found and many of these only with few species.
Little is known about the biology of these beetles. Most feed on fungi, others are carnivores and eat small arthropods such as bark beetles.

Systematics and taxonomy

Up to 9 tribes are accepted by various authors; others, however, synonymize some of these. Formerly, many additional tribes were recognized, but the Synchitini, for example, are today generally held to include a number of these tribes, and are even sometimes merged into the Colydiini.
The tribes are:
  • Acropini
  • Adimerini
  • Colydiini Billberg, 1820
  • Gempylodini
  • Nematidiini
  • Orthocerini Blanchard, 1845
  • Rhagoderini
  • Rhopalocerini
  • Synchitini Erichson, 1845
Delimitation of the Colydiinae against the other lineages of Zopheridae is usually unproblematic. The only significant case of dispute may be the Pycnomerini, which is a small lineage of Zopheridae incertae sedis and was formerly considered an independent family like the "Colydiidae". That treatment is almost certainly wrong, but whether these beetles should be placed in Zopheridae as an additional subfamily Pycnomerinae, or treated as tribe Pycnomerini - and if the latter, whether they are better included in the Colydiinae or the Zopherinae - is still disputed.

Selected genera

Genera of cylindrical bark beetles include:Ablabus Broun, 1880AcolobicusAcolophoides Slipinski & Lawrence, 1997Acolophus Sharp, 1885Acostonotus Slipinski & Lawrence, 1997Acropis Burmeister, 1840AfrorthocerusAllobitoma Broun, 1921AlluauditomaAnosyanaAntilissus Sharp, 1879AscommaAsproteraAsynchitaAtyscusAulonium Erichson, 1845BhutaniaBitoma Herbst, 1793Bolcocius Dajoz, 1977Bulasconotus Ślipiński & Lawrence, 1997BupalaCacotarphiusCaprodes Pascoe, 1863CatolaemusCebia Pascoe, 1863Cerchanotus Erichson, 1845Chorasus Sharp, 1882 Cicablabus Slipinski & Lawrence, 1997CiconesCiconissus Broun, 1893 Colobicones Grouvelle, 1918Colobicus Latreille, 1807Colydium Fabricius, 1792ColydodesCorticus Germar, 1824Coxelus Dejean, 1821DechomusDenophloeusDiodesma Latreille, 1829Diplagia Reitter, 1882DiplotomaDitomoideaEmilkaEndeitoma Sharp, 1894EndocoxelusEndophloeus Dejean, 1834Enhypnon Carter, 1919Epistranodes Slipinski & Lawrence, 1997Epistranus Sharp, 1878ErylusEuciconesEudesmaEulachusFaecula Slipinski & Lawrence, 1997FeneriviaGempylodes Pascoe 1863Glenentela Broun, 1893Glyphocryptus Sharp, Helioctamenus Schaufuss, 1882Heterargus Sharp, 1886 HolopleuridiaHyberisHystriconesIsotarphiusKanantsiaLabrotrichusLangelandia Aubé, 1843Lascobitoma Slipinski & Lawrence, 1997Lasconotus Erichson, 1845LascotonusLascotrichusLastrema Reitter, 1882LinophloeusLobogestoriaLobomesa Slipinski & Lawrence, 1997Lyreus Aubé, 1861MadaconesMadadesiaMadenphloeusMamakiusMegataphrusMicroprius Fairmaire, 1868MicrosicusMnionychusMonoedus Horn, 1882 Munaria Reitter, 1882Namunaria Reitter, 1882 Nematidium Erichson, 1845Neotrichus Sharp, 1885Niphopelta Reitter, 1882Norix Broun, 1893Nosodomodes Reitter, 1922Notocoxelus Ślipiński & Lawrence, 1997Orthocerus Latreille, 1796 PahaParatarphiusParyphusPharaxPhloeodalisPhloeonemusPhormesaPhorminx Carter & Zeck, 1937PhreatusPriolomopsisPriolomusPristoderus Hope, 1840 Prosteca Wollaston, 1860Pseudendestes Lawrence, 1980PseudocorticusPseudotarphiusRechodesRhagodera Mannerheim 1843Rhopalocerus Redtenbacher, 1842Rytinotus Broun, 1880SallachiusSassakaSechellotomaSprecodesSynagathis Carter & Zeck, 1937Syncalus Sharp, 1876 Synchita Hellwig, 1792Tarphiablabus Ślipiński & Lawrence, 1997Tarphiomimus Wollaston, 1873TarphiosomaTarphius Erichson, 1845Tentablabus Slipinski & Lawrence, 1997Todima Grouvelle, 1893Todimopsis Ślipiński & Lawrence, 1997Trachypholis Erichson, 1845TrigonophloeusXylolaemus Redtenbacher, 1858Zanclea
The genera Pycnomerodes, Pycnomerus and Rhizonium are sometimes included in the Colydiinae too. Other authors consider the latter incertae sedis among the Tenebrionoidea; for the former two, see [|above]. The species Xylolaemus sakhnovi was described in 2014 from a fossil preserved in Baltic amber, which dates to the Middle Eocene. This was the first species of Xylolaemus described from the fossil record. The extinct genus Paleoendeitoma, belonging to the extant tribe Synchitini is known from the Cenomanian aged Burmese amber of Myanmar.