Pentatominae


Pentatominae is a subfamily of Pentatomidae, a family of shield bugs. This subfamily is the largest one within the Pentatomidae, having 4937 species classified in 938 genera. Species in this subfamily are phytophages and several of them are considered agricultural pests. Some invasive pentatomines such as Halyomorpha halys and Bagrada hilaris have been considered household pests. Higher systematics of the group have been revised by Rider et al.

Tribes and genera

Tribes within this subfamily include:
  • Aelini Douglas & Scott, 1865
  1. Aelia Fabricius, 1803
  2. Aeliopsis Bergevin, 1931
  3. Neottiglossa Kirby, 1837
  • Aeptini Stål, 1871
  1. Aeliosoma Baehr, 1985
  2. Aeptus Dallas, 1851
  3. Caridillus Bergroth, 1909
  4. Eribotes Stål, 1867
  5. Hillieria Distant, 1910
  6. Mariomella Faúndez & Rider, 2019
  7. Menestheus Stål, 1867
  8. Paramenestheus Breddin, 1900
  9. Pseudaelia Distant, 1910
  • Aeschrocorini Distant, 1902
  1. Aeschrocoris Bergroth, 1887
  2. Aeschrus Spinola, 1850
  3. Chraesus Cachan, 1952
  4. Geomorpha Bergroth, 1893
  5. Risbecella Schouteden, 1942
  6. Scylax Distant, 1887
  7. Tyoma Spinola, 1850
  8. Tyomana Miller, 1952
  • Agaeini Cachan, 1952
  1. Agaeus Dallas, 1851
  • Agonoscelidini Atkinson, 1888
  1. Agonoscelis Spinola, 1837
  • Amyntorini Distant, 1902
  1. Ambohicorypha Cachan, 1952
  2. Belopis Distant, 1879
  3. Bolaca Walker, 1867
  4. Halyabbas Distant, 1900
  5. Madecorypha Cachan, 1952
  6. Sennertus Distant, 1900
  1. Holcogaster Fieber, 1860
  • Axiagastini Atkinson, 1888
  1. Acesines Stål, 1876
  2. Axiagastus Dallas, 1851
  3. Indrapura China, 1928
  4. Oncotropis Stål, 1871
  1. Bathycoelia Amyot & Serville, 1843
  • Cappaeini Atkinson, 1888
  1. Adelolcus Bergroth, 1893
  2. Benia Schouteden, 1916
  3. Boerias Kirkaldy, 1909
  4. Cappaea Ellenrieder, 1862
  5. Caura Stål, 1865
  6. Cauromorpha Jeannel, 1913
  7. Erlangerella Schouteden, 1905
  8. Halycorypha Jeannel, 1913
  9. Halydicoris Jeannel, 1913
  10. Halyoides Cachan, 1952
  11. Halyomorpha Mayr, 1864
  12. Homalogonia Jakovlev, 1876
  13. Hymenomaga Karsch, 1892
  14. Lerida Karsch, 1894
  15. Leridella Jeannel, 1913
  16. Lokaia Linnavuori, 1975
  17. Mabusana Distant, 1912
  18. Massocephalus Dallas, 1851
  19. Paralerida Linnavuori, 1982
  20. Prytanicoris Gross, 1978
  21. Tolumnia Stål, 1867
  22. Tripanda Berg, 1899
  23. Tropicorypha Mayr, 1864
  24. Veterna Stål, 1865
  1. Arocera Spinola, 1837
  2. Boea Walker, 1867
  3. Catacanthus Spinola, 1837
  4. Chalcocoris Dallas, 1851
  5. Rhyssocephala Rider, 1992
  6. Runibia Stål, 1861
  7. Vitruvius Distant, 1901
  8. Vulsirea Spinola, 1837
  • Caystrini Ahmad & Afzal, 1979
  1. Caystrus Stål, 1861
  2. Cephaloplatus White, 1842
  3. Heissocoris Ahmad & Afzal, 1979
  4. Hippotiscus Bergroth, 1906
  5. Kyrtalus Van Duzee, 1929
  6. Lyalocoris Zahid, 2006
  7. Mcdonaldiana Zahid, 2006
  8. Neodius
  9. Neoparodius Zahid, 2006
  10. Neoriazocoris Zahid, 2006
  11. Paramecocoris Stål, 1853
  12. Parodius Distant, 1900
  13. Praetextatus Distant, 1901
  14. Riazocoris Ahmad & Afzal, 1979
  15. Valescus Distant, 1901
  • Chlorocorini Rider, Greve, Schwertner & Grazia, 2018
  1. Arvelius Spinola, 1837
  2. Chlorocoris Spinola, 1837
  3. Chloropepla Stål, 1867
  4. Eludocoris Thomas, 1992
  5. Fecelia Stål, 1872
  6. Mayrinia Horváth, 1925
  7. Rhyncholepta Bergroth, 1911
  • Coquereliini Cachan, 1952
  1. Cleoqueria Cachan, 1952
  2. Coquerelia Signoret, 1861
  3. Coquerelidea Reuter, 1887
  4. Neocoquerelidea Cachan, 1952
  • Degonetini Azim & Shafee, 1984
  1. Degonetus Distant, 1902
  1. Ambiorix Stål, 1876
  2. Diplostira Dallas, 1851
  • Diploxyini Atkinson, 1888
  1. Acoloba Spinola, 1850
  2. Aeladria Cachan, 1952
  3. Diploxys Amyot & Serville, 1843
  • Eurysaspidini Atkinson, 1888
  1. Eurysaspis Signoret, 1851
  2. Flaminia Stål, 1865
  • Eysarcorini Mulsant & Rey, 1866
  1. Adria Stål, 1876
  2. Aspavia Stål, 1865
  3. Cachanocoris Rider, 1998
  4. Carbula Stål, 1864
  5. Corisseura Cachan, 1952
  6. Cratonotus Distant, 1879
  7. Durmia Stål, 1865
  8. Dymantiscus Hsiao, 1981
  9. Empiesta Linnavuori, 1982
  10. Eysarcoris Hahn, 1834
  11. Hermolaus Distant, 1902
  12. Mulungua Linnavuori, 1982
  13. Pseudolerida Schouteden, 1958
  14. Sepontia Stål, 1865
  15. Sepontiella Miyamoto, 1990
  16. Spermatodes Bergroth, 1914
  17. Stagonomus Gorski, 1852
  • Halyini Amyot & Serville, 1843 - selected genera:
  • * Brochymena Amyot & Serville, 1843
  • Hoplistoderini Atkinson, 1888
  1. Alcimocoris Bergroth, 1891
  2. Glottaspis Bergroth, 1918
  3. Hoplistodera Westwood, 1837
  4. Paracritheus Bergroth, 1891
  5. Paralcimocoris Zhang & Lin, 1988
  6. Stachyomia Stål, 1871
  7. Taurodes Dallas, 1851
  • Lestonocorini Ahmad & Mohammad, 1980
  1. Dardjilingia Yang, 1936
  2. Gynenica Dallas, 1851
  3. Lestonocoris Ahmad & Mohammad, 1980
  4. Neogynenica Yang, 1935
  5. Umgababa Leston, 1953
  1. Mecidea Dallas, 1851
  • Memmiini Cachan, 1952
  1. Memmia Stål, 1865
  • Menidini Atkinson, 1888
  1. Actuarius Distant, 1901
  2. Aegaleus Stål, 1876
  3. Amphimachus Stål, 1867
  4. Anchesmus Stål, 1876
  5. Apines Dallas, 1851
  6. Aspideurus Signoret, 1880
  7. Banya Schouteden, 1916
  8. Brachycoris Stål, 1871
  9. Carvalhocoris Leston, 1953
  10. Cresphontes Stål, 1867
  11. Dabessus Distant, 1902
  12. Decellella Schouteden, 1964
  13. Desertomenida Kiritshenko, 1914
  14. Dunnius Distant, 1902
  15. Elanela Rolston, 1980
  16. Eurymenida Bergroth, 1917
  17. Gwea Schouteden, 1958
  18. Keleacoris Rider & Rolston, 1995
  19. Lathraedoeus Breddin, 1900
  20. Menida Motschulsky, 1861
  21. Neostrachia Saunders, 1877
  22. Phavorinus Distant, 1902
  23. Rio Kirkaldy, 1909
  24. Saceseurus Breddin, 1900
  25. Sciomenida Gross, 1976
  26. Solenocoris Oshanin, 1922
  27. Udonga Distant, 1921
  • Myrocheini Stål, 1871
  1. Aednulus Breddin, 1901
  2. Aednus Dallas, 1851
  3. Arniscus Distant, 1899
  4. Delegorguella Spinola, 1850
  5. Dictyotus Dallas, 1851
  6. Dictytotus Dallas, 1851
  7. Dissocolpus Bergroth, 1906
  8. Dollingiana Ahmad & Kamaluddin, 1986
  9. Dymantis Stål, 1861
  10. Ennius Stål, 1861
  11. Erachtheus Stål, 1861
  12. Humria Linnavuori, 1975
  13. Jitka Kment, 2015
  14. Laprius Stål, 1861
  15. Lodosia Ahmad & Önder, 1996
  16. Munshiana Ahmad & Kamaluddin, 1987
  17. Myrochea Amyot & Serville, 1843
  18. Neococalus Bergroth, 1891
  19. Neodorpius Ahmad & Afzal, 1989
  20. Paradictyotus Gross, 1975
  21. Pretorius Distant, 1898
  22. Stysicoris Ahmad & Kamaluddin, 1985
  23. Tholosanus Distant, 1899
  24. Utheria Gross, 1975
  • Nealeriini Cachan, 1952
  1. Nealeria Bergroth, 1893
  2. Paraleria Reuter, 1887
  • Nezarini Atkinson, 1888 - selected genera:
  • * Nezara Amyot et Serville, 1843
  • * Palomena Mulsant et Rey, 1866
  • Opsitomini Cachan, 1952
  1. Opsitoma Cachan, 1952
  • Pentamyrmexini Rider & Brailovsky, 2014
  • * Pentamyrmex Rider & Brailovsky, 2014
  • Pentatomini Leach, 1815 - selected genera:
  • * Pentatoma Olivier, 1789
  • Phricodini Cachan, 1952
  1. Phricodus Spinola, 1839
  • Piezodorini Atkinson, 1888
  1. Anaximenes Stål, 1876
  2. Chaubattiana Distant, 1912
  3. Pausias Jakovlev, 1905
  4. Piezodorus Fieber, 1860
  • Procleticini Pennington, 1920
  1. Aleixus McDonald, 1981
  2. Brepholoxa Van Duzee, 1904
  3. Dendrocoris Bergroth, 1891
  4. Lobepomis Berg, 1891
  5. Neoderoploa Pennington, 1922
  6. Odmalea Bergroth, 1914
  7. Parodmalea Rider, 1994
  8. Procleticus Berg, 1891
  9. Pseudobebaeus Distant, 1911
  10. Terania Pirán, 1963
  11. Thoreyella Spinola, 1850
  1. Agathocles Stål, 1876
  2. Amasenus Stål, 1863
  3. Amatembuna Distant, 1910
  4. Critheus Stål, 1867
  5. Exithemus Distant, 1902
  6. Rolstoniellus Rider, 1997
  1. Anoano Cachan, 1952
  2. Nenes Kment, 2015
  3. Tricompastes Cachan, 1952
  4. Triplatyx Horváth, 1904

Genera ''[incertae sedis]''

  1. Amphidexius Bergroth, 1918
  2. Bachesua Gross, 1975
  3. Birna McDonald, 2006
  4. Brizocoris Gross, 1975
  5. Buthumka Gross, 1975
  6. Cooperocoris Gross, 1975
  7. Dysnoetus Bergroth, 1918
  8. Eremophilacoris Gross, 1975
  9. Gadarscama Reuter, 1887
  10. Ippatha Distant, 1910
  11. Kitsonia Gross, 1975
  12. Kitsoniocoris Rider, 1998
  13. Kumbutha Distant, 1910
  14. Lakhonia Yang, 1936
  15. Macrocarenoides Gross, 1975
  16. Macrocarenus Stål, 1873
  17. Mcphersonarcys Thomas, 2012
  18. Muscanda Walker, 1868
  19. Ochisme Kirkaldy, 1904
  20. Ooldeon Gross, 1975
  21. Patanius Rolston, 1987
  22. Pirricoris Gross, 1975
  23. Poecilotoma Dallas, 1851
  24. Riaziana Hasan, Afzal & Ahmad, 1989
  25. Senectius Rolston, 1987
  26. Tepperocoris Gross, 1975
  27. Thryptomenecoris Gross, 1975
  28. Trachyops Dallas, 1851
  29. Turrubulana Distant, 1910
  30. Xiengia Distant, 1921
  31. Zouicoris Zheng, 1986

Notable species