Groundhog


The groundhog, also known as the woodchuck, is a rodent of the family Sciuridae, belonging to the group of large ground squirrels known as marmots.
A lowland creature of North America, it is found through much of the Eastern United States, across Canada and into Alaska.
It was given its scientific name as Mus monax by Carl Linnaeus in 1758, based on a description of the animal by George Edwards, published in 1743.
The groundhog, being a lowland animal, is exceptional among marmots. Other marmots, such as the yellow-bellied and hoary marmots, live in rocky and mountainous areas. Groundhogs are considered one of the most solitary of marmot species. They live in aggregations, and their social organization and long-term pair bonds varies across populations. The groundhog's male and female interactions are usually limited to the mating season and copulation. However, certain populations of groundhogs have been observed to form long-term adult male-female association throughout the year, and often from year to year.
The groundhog is an important contributor to the maintenance of healthy soil in woodlands and plains; as such, the species is considered a crucial habitat engineer. The groundhog is an extremely intelligent animal, forming complex social networks and kinship with its young; it is capable of understanding social behavior, communicating threats through whistling, and working cooperatively to accomplish tasks such as burrowing.

Etymology

Common names for the groundhog include chuck, wood-chuck, groundpig, whistle-pig, whistler, thickwood badger, Canada marmot, monax, moonack, weenusk, red monk, land beaver and, among French Canadians in eastern Canada, siffleux. The name "thickwood badger" was given in the Northwest to distinguish the animal from the prairie badger. Monax is an Algonquian name of the woodchuck, which means "digger". Young groundhogs may be called chucklings.
The etymology of the name woodchuck is unrelated to wood nor any sense of chucking. It stems from an Algonquian name for the animal, wuchak. The similarity between the words has led to the popular tongue-twister:

Description

The groundhog is the largest sciurid in its geographical range, excluding its presence in British Columbia where its range may be comparable to that of its somewhat larger cousin, the hoary marmot. Adults may measure from in total length, including a tail of. Weights of adult groundhogs typically fall between.
Male groundhogs are slightly larger than females on average and, like all marmots, they are considerably heavier during autumn than when they emerge from hibernation in spring. Adult males average year-around weight, with spring to fall average weights of while females average, with spring to fall averages of. Seasonal weight changes reflect circannual deposition and use of fat. Groundhogs attain progressively higher weights each year for the first two or three years, after which weight plateaus.
Groundhogs have four incisors, which grow per week. Constant usage wears them down by about that much each week. Unlike the incisors of many other rodents, the incisors of groundhogs are white to ivory-white. Groundhogs are well-adapted for digging, with powerful, short legs and broad, long claws. The groundhog's tail is shorter than that of other Sciuridae—only about one-fourth of body length.

Distribution and habitat

The groundhog dwells in lowland habitats, unlike other marmots that live in rocky and mountainous areas. Marmota monax has a wide geographic range. The groundhog prefers open country and the edges of woodland, being rarely found far from a burrow entrance. It can typically be found in small woodlots, low-elevation forests, fields and pastures, and hedgerows. It constructs dens in well-drained soil, and most groundhogs have summer and winter dens. Human activity has increased food access and abundance, allowing M. monax to thrive.

Behavior

W.J. Schoonmaker reports that groundhogs may hide when they see, smell, or hear an observer. Marmot researcher Ken Armitage states that the social biology of the groundhog is poorly studied.
Despite their heavy-bodied appearance, groundhogs are accomplished swimmers and occasionally climb trees when escaping predators or when they want to survey their surroundings. They prefer to retreat to their burrows when threatened; if the burrow is invaded, the groundhog tenaciously defends itself with its two large incisors and front claws. Groundhogs are generally agonistic and territorial toward each other and may skirmish to establish dominance.
Outside their burrow, individuals are alert when not actively feeding. It is common to see one or more nearly motionless individuals standing erect on their hind feet watching for danger. When alarmed, they use a high-pitched whistle to warn the rest of the colony, hence the name "whistle-pig". Groundhogs may squeal when fighting, seriously injured, or caught by a predator. Other vocalizations include low barks and a sound produced by grinding their teeth. David P. Barash wrote that he witnessed only two occasions of upright play-fighting among woodchucks and that the upright posture of play-fighting involves sustained physical contact between individuals that may require a degree of social tolerance virtually unknown in M. monax. Alternatively, upright play-fighting may be a part of the woodchuck's behavioral repertoire that rarely is shown because of physical spacing and/or low social tolerance.

Diet

Mostly herbivorous, groundhogs eat primarily wild grasses and other vegetation, including berries, bark, leaves, and agricultural crops, when available. In early spring, dandelion and coltsfoot are important groundhog food items. Some additional foods include sheep sorrel, timothy-grass, buttercup, persicaria, agrimony, red and black raspberries, mulberries, buckwheat, plantain, wild lettuce, alfalfa, and all varieties of clover. Groundhogs also occasionally eat small animals, such as grubs, grasshoppers, snails, and even bird eggs and baby birds, but are not as omnivorous as many other Sciuridae.
An adult groundhog can eat more than of vegetation daily. In early June, woodchucks' metabolism slows, and while their food intake decreases, their weight increases by as much as 100% as they produce fat deposits to sustain them during hibernation and late winter. Instead of storing food, groundhogs stuff themselves to survive the winter without eating. Thought not to drink water, groundhogs are reported to obtain needed liquids from the juices of edible plants, aided by their sprinkling with rain or dew.

Burrows

Groundhogs are excellent burrowers, using burrows for sleeping, rearing young, and hibernating. Groundhog burrows usually have two to five entrances, providing groundhogs their primary means of escape from predators. The volume of earth removed from groundhog burrows in one study averaged per den. The longest burrow measured in addition to two short side galleries.
Though groundhogs are the most solitary of the marmots, several individuals may occupy the same burrow.
Burrows can pose a serious threat to agricultural and residential development by damaging farm machinery and even undermining building foundations. In a June 7, 2009, Humane Society of the United States article, "How to Humanely Chuck a Woodchuck Out of Your Yard" by John Griffin, director of Humane Wildlife Services, stated you would have to have a lot of woodchucks working over a lot of years to create tunnel systems that would pose any risk to a structure.
The burrow is used for safety, retreat in bad weather, hibernating, sleeping, mating, and nursery. In addition to the nest, there is an excrement chamber. The hibernation or nest chamber is lined with dead leaves and dried grasses. The nest chamber may be about twenty inches to three feet below ground surface. It is about wide and high. There are typically two burrow openings or holes. One is the main entrance, the other a spy hole. Description of the length of the burrow often includes side galleries. Excluding side galleries, Schoonmaker reports the longest was, and the average length of eleven dens was. W. H. Fisher investigated nine burrows, finding the deepest point down. The longest, including side galleries, was. Numbers of burrows per individual groundhog decrease with urbanization.
Bachman mentioned when young groundhogs are a few months old, they prepare for separation, digging a number of holes in the area of their early home. Some of these holes were only a few feet deep and never occupied, but the numerous burrows gave the impression that groundhogs live in communities.
Abandoned groundhog burrows benefit many other species by providing shelter. They are used by cottontail rabbits, raccoons, foxes, river otters, eastern chipmunks, and a wide variety of small mammals, snakes, and birds.

Hibernation

Groundhogs are one of the few species that enter into true hibernation, and often build a separate "winter burrow" for this purpose. This burrow is usually in a wooded or brushy area and is dug below the frost line and remains at a stable temperature well above freezing during the winter months. In most areas, groundhogs hibernate from October to March or April, but in more temperate areas, they may hibernate as little as three months. Groundhogs hibernate longer in northern latitudes than southern latitudes. To survive the winter, they are at their maximum weight shortly before entering hibernation. When the groundhog enters hibernation, there is a drop in body temperature to as low as, heart rate falls to 4–10 beats per minute and breathing rate falls to one breath every six minutes. During hibernation, they experience periods of torpor and arousal. Hibernating woodchucks lose as much as half their body weight by February. They emerge from hibernation with some remaining body fat to live on until the warmer spring weather produces abundant plant materials for food. Males emerge from hibernation before females. Groundhogs are mostly diurnal and are often active early in the morning or late afternoon.