Urocyclidae


Urocyclidae is a family of air-breathing land snails, semi-slugs and land slugs, terrestrial pulmonate gastropod mollusks in the superfamily Helicarionoidea.
Some species within this family make and use love darts before mating.

Distribution

The Urocyclidae are distributed in the Afrotropical realm, including Madagascar, the Comores and the islands of the Gulf of Guinea.

Taxonomy

The family Urocyclidae consists of 3 subfamilies:
  • subfamily Rhysotininae Schileyko, 2002
  • subfamily Sheldoniinae Connolly, 1925 - synonyms: Peltatinae Godwin-Austen, 1912; Trochonanininae Connolly, 1912; Trochozonitinae Iredale, 1914; Ledoulxiinae Pilsbry, 1919; Gymnarionidae Von Mol, 1970; Rhysotinidae Schileyko, 2002; Zonitarionini Schileyko, 2002; Acantharionini Schileyko, 2002
  • subfamily Urocyclinae Simroth, 1889
  • * tribe Urocyclini Simroth, 1889 - synonyms: Atoxonini Schileyko, 2002; Buettneriini Schileyko, 2002
  • * tribe Dendrolimacini Van Goethem, 1977
  • * tribe Leptichnini Van Goethem, 1977
  • * tribe Upembellini Van Goethem, 1977

Genera

The following genera are recognised in the family Urocyclidae:
; unsorted
  • Koruella Pickford, 2009
;Subfamily RhysotininaeRhysotina Ancey, 1887
;Subfamily SheldoniinaeAcantharion Binder & Tillier, 1985Africarion Godwin-Austen, 1883Amatarion Van Mol, 1970Angustivestis Pilsbry, 1919Aspidelus Morelet, 1884Aspidotomium Degner, 1932Belonarion Pilsbry, 1919Bloyetia Bourguignat, 1890Camerunarion Van Mol, 1970Carinazingis van Bruggen & de Winter, 1990Centrafricarion Van Mol, 1970Chlamydarion Van Mol, 1968Colparion Laidlaw, 1938Degneria Verdcourt, 1956Entagaricus Pilsbry, 1919Estria Poirier, 1887Falloonella Preston, 1914Fuellebornia Verdcourt, 1998Granularion Germain, 1912Gymnarion Pilsbry, 1919
  • Hamya Bourguignat, 1885 Kerkophorus Godwin-Austen, 1912Lacrimarion Connolly, 1939Malagarion Tillier, 1979Mesafricarion Pilsbry, 1919Microkerkus Godwin-Austen, 1912Montanobloyetia Verdcourt, 1961Morrumbalia Verdcourt, 1998Paragranularion Van Mol, 1970Percivalia Preston, 1914Plicatonanina Verdcourt, 1961Principicochlea D. Holyoak & G. Holyoak, 2020Principitrochoidea D. Holyoak & G. Holyoak, 2020Ptilototheca Herbert, 2016Rhopalogonium Degner, 1932Selatodryas Herbert, 2017Senegalarion Van Mol, 1970Sheldonia Ancey, 1887 - the type genus of the subfamily SheldoniinaeSjostedtina Verdcourt, 1961Sylvarion Van Mol, 1970Thapsia Albers, 1860Thielarion Van Mol, 1970Thomithapsia D. Holyoak & G. Holyoak, 2020Thomitrochoidea D. Holyoak & G. Holyoak, 2020Tresia Van Goethem, 1975Trochonanina Mousson, 1869Trochozonites Pfeffer, 1883Tropidocochlion Verdcourt, 1998Verdcourtia Van Mol, 1970Verrucarion Van Mol, 1970Zingis E. von Martens, 1878Zonitarion Pfeffer, 1883
; Subfamily Urocyclinae
Tribe DendrolimaciniDendrolimax Heynemann, 1868 - the type genus of the tribe Dendrolimacini
Tribe LeptichniniLeptichnus Simroth, 1896 - the type genus of the tribe Leptichnini
Tribe UpembelliniLeptichnoides Van Goethem, 1975Upembella Van Goethem, 1969 - the type genus of the tribe Upembellini
tribe UrocycliniAnisotoxon Van Goethem, 1975Atoxon Simroth, 1888Atoxonoides Van Goethem, 1973Atrichotoxon Simroth, 1910Buettneria Simroth, 1910 Bukobia Simroth, 1896Elisolimax Cockerell, 1893Emphysetes Verdcourt, 2003Microcyclus Simroth, 1896Nupnus Van Goethem, 1975Pembatoxon Van Goethem, 1975Phaneroporus Simroth, 1888Polytoxon Simroth, 1897Pseudatoxon Van Goethem, 1975Tanzalimax Rowson, Paustian & Van Goethem, 2017Trichotoxon Simroth, 1888Udzungwalimax Rowson, Paustian & Van Goethem, 2017Urocyclus Gray, 1864 - the type genus of the family Urocyclidae

Cladogram

The following cladogram shows the phylogenic relationship of this family to other families in the limacoid clade: