Umbra (fish)

Umbra is a genus of mudminnows native to Europe and North America. It is the only extant member of the family Umbridae.
Phylogenetic evidence suggests the North American and European clades split between the Late Cretaceous or early Paleogene, making this a rather ancient group.

Species

The currently recognized species in this genus are:Umbra krameri Walbaum, 1792 Umbra limi Umbra pygmaea
The following fossil species are also known:

†Umbra euronota Schwarzhans et al., 2024 - Middle Miocene of Ukraine
†Umbra irtyshensis Sytchevskaya, 1968
†Umbra longidorsalis Böhme, 2004 - Early Miocene of the Czech Republic
†Umbra oderiensis Oberhlová, 1978 - Late Oligocene/Early Miocene of the Czech Republic
†Umbra perpusilla - Middle Miocene of Germany
†Umbra praekrameri Weinfurter, 1950 - Late Miocene of Austria
†Umbra prochazkai Oberhlová, 1978 - Late Oligocene of the Czech Republic

Indeterminate Umbra scales are known from the Late Eocene/Early Oligocene of the Czech Republic.