Sticherus


Sticherus is a genus of about 80–90 species of fern.
In Australia they are commonly referred to as shield ferns. In the United States they are commonly referred to as umbrella ferns.
The fronds are branched at least once with each pair of branches spreading out, forming an umbrella like layer of foliage. Fronds are mostly hairless and often glaucous beneath the frond.

Phylogeny

The following species are recognised as of February 2023.
Phylogeny of Sticherus
Other species include:S. albus Gonzales 2011S. antillensis Gonzales 2011S. arachnoideus Østerg. & Øllgaard 2001S. aurantiacus Østerg. & Øllgaard 2001S. blepharolepis Ching 1940S. bolanicus Copeland 1941S. boliviensis Gonzales 2011S. brassii Copeland 1941S. brevitomentosus Østerg. & Øllgaard 2001S. chocoensis Gonzales 2011S. cubensis Gonzales 2011S. cundinamarcensis Nakai 1950S. decurrens Gonzales 2011S. erectus Copeland 1941S. farinosus Ching 1940S. ferrugineus Gonzales 2011S. fulvus Ching 1940S. fuscus Gonzales 2011S. gnidioides Nakai 1950S. gracilis Copeland 1947S. hastulatus Nakai 1950S. hirtus Ching 1940S. hispaniolensis GonzalesS. hispidus Copeland 1947S. holttumii Lima & Salino 2018S. hooglandii Perrie 2018S. impressus Parris 1992S. inflexus Pichi Sermolli 1972S. interjectus Gonzales 2011S. intermedius Chrysler 1944S. jacha Gonzales 2011S. jamaicensis Nakai 1950S. lanosus Gonzales 2011S. lanuginosus Nakai 1950S. lechleri Nakai 1950S. ×leonis Gonzales 2011S. lepidotus R.A.Rodr. & Ponce 2007S. litoralis Nakai 1950S. lobatus Wakef. 1943 S. loheri Copeland 1947S. longipinnatus Ching 1940S. maritimus Nakai 1950S. melanoblastus Østerg. & Øllgaard 2001S. milnei Ching 1940S. montaguei Nakai 1950S. moyobambensis Gonzales 2011S. nervatus Gonzales 2011S. nigropaleaceus Prado & Lellinger 1996S. nudus Nakai 1950S. oceanicus Ching 1940S. orthocladus Chrysler 1944S. ovatus Gonzales 2011S. pallescens Vareschi 1969S. penniger Copeland 1947S. peruvianus Smith, Kessler & Gonzales 1999S. pruinosus Ching 1940S. ×pseudobifidus Gonzales 2011S. pseudoscandens Copeland 1941S. pteridellus Copeland 1947S. pulcher Copeland 1941S. quadripartitus Ching 1940S. reflexipinnulus Copeland 1941S. remotus Chrysler 1944S. revolutus Ching 1940S. rubiginosus Nakai 1950S. rufus Gonzales 2011S. salinoi Lima 2018S. simplex Ching 1940S. squamosus Gonzales 2011S. squamulosus Nakai 1950S. strictissimus Copeland 1947S. ×subremotus Gonzales 2011S. tepuiensis Smith 1990S. tomentosus Smith 1990S. umbraculifer Ching 1940S. underwoodianus Nakai 1950S. velatus Copeland 1947S. venosus Copeland 1941S. vestitus Ching 1940