Sam W. Heads


Sam W. Heads is a British palaeontologist, a Fellow of the Linnean Society of London, a Fellow of the Royal Entomological Society, as well as a former Officer and Editor-in-Chief at the Orthopterists' Society.
He received his training at the University of Portsmouth where he was awarded a B.Sc. in Palaeobiology and Evolution in 2004 and a Ph.D. in Entomology in 2009. Heads is an authority on the taxonomy, systematics and palaeontology of Orthoptera. He is based at the Illinois Natural History Survey, University of Illinois at Urbana-Champaign.

Select publications

  • Heads, S.W. & Wang, Y. 2013. First fossil record of Melanoplus differentialis. Entomological News 123: 33–37.
  • Heads, S.W., Taylor, S.J. & Krejca, J.K. 2013. First record of Scapteriscus abbreviatus from Belize. Entomological News 123: 241–244.
  • Hollier, J. & Heads, S.W. 2014. The type specimens of Orthoptera described by American entomologists in the Muséum d’histoire naturelle de Genève. Revue suisse de Zoology 121: 63–76.
  • Barling, N., Heads, S.W. & Martill, D.M. 2013. A new parasitoid wasp from the Lower Cretaceous Crato Formation of Brazil: the first Mesozoic Pteromalidae. Cretaceous Research 45: 258–264,
  • Hollier, J., Bruckner, H. & Heads, S.W. 2013. An annotated list of the Orthoptera species described by Henri de Saussure, with an account of the primary type material housed in the Muséum d’histoire naturelle de Genève, Part 5: Grylloidea. Revue suisse de Zoology 120: 445–535.
  • Jepson, J.E., Heads, S.W., Makarkin, V.N. & Ren, D. 2013. New fossil mantidflies from the Mesozoic of northeastern China. Palaeontology 56: 603–613.
  • Heads, S.W., Penney, D. & Green, D.I. 2012. A new species of the cricket genus Proanaxipha in Miocene amber from the Dominican Republic. ZooKeys 229: 111–118.
  • Pérez-de la Fuente, R., Heads, S.W., Hinojosa-Díaz, I.A. & Engel, M.S. 2012. The first record of Protogryllinae from the Jurassic of India. Journal of the Kansas Entomological Society 85: 53–58.
  • Hollier, J. & Heads, S.W. 2012. An annotated list of the Orthoptera species described by Henri de Saussure, with an account of the primary type material housed in the Muséum d’histoire naturelle de Genève, Part 1: Tridactyloidea. Revue suisse de Zoology 119: 149–160.
  • Rodríguez, F. & Heads, S.W. 2012. New mole crickets of the genus Scapteriscus Scudder from Colombia. Zootaxa 3282: 61–68.
  • Heads, S.W. & Taylor, S.J. 2012. A new species of Ripipteryx from Belize with a key to the species of the Scrofulosa Group. ZooKeys 169: 1–8.
  • Heads, S.W. & Leuzinger, L. 2011. On the placement of the Cretaceous orthopteran Brauckmannia groeningae from Brazil, with notes on the relationships of Schizodactylidae. ZooKeys 77: 17–30.
  • Heads, S.W. 2010. New Tridactyloidea in Miocene amber from the Dominican Republic. Annales de la Société Entomologique de France 46: 204–210.
  • Heads, S.W. 2010. A new species of Ripipteryx from the Ecuadorian Andes. Zootaxa 2476: 23–29.
  • Heads, S.W. 2010. The first fossil spider cricket : 20 million years of troglobiomorphosis or exaptation in the dark? Zoological Journal of the Linnean Society 158: 56–65.
  • Heads, S.W. & Maehr, M.D. 2009. Proposed conservation of the name Gastrimargus Saussure, 1884 threatened by an unused senior homonym. Zootaxa 2268: 65–68.
  • Heads, S.W. 2009. A new pygmy mole cricket in Cretaceous amber from Burma. Denisia 26: 75–82.
  • Heads, S.W. 2009. New pygmy grasshoppers in Miocene amber from the Dominican Republic. Denisia 26: 69–74.
  • Delclòs, X., Nel, A., Azar, D. Bechly, G., Dunlop, J.A., Engel, M.S. & Heads, S.W. 2008. The enigmatic Mesozoic insect taxon Chresmodidae : new palaeobiological and phylogenetic data, with the description of a new species from the Lower Cretaceous of Brazil. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 247: 353–381.
  • Heads, S.W., Martill, D.M. & Loveridge, R.F. 2008. Palaeoentomological paradise: the Cretaceous Crato Formation of Brazil. Antenna 32: 91–98.
  • Heads, S.W. & Chesmore, D. 2008. New records of the slender ground-hopper Tetrix subulata from Yorkshire. British Journal of Entomology & Natural History 21: 243–246.
  • Heads, S.W. & Lakin, C. 2008. A new katydid of the genus Phoebolampta from St Maarten. Polskie Pismo Entomologiczne 77: 256–266.
  • Heads, S.W. 2008. A new species of Yuripopovia from the Early Cretaceous of the Isle of Wight. British Journal of Entomology & Natural History 21: 247–254.
  • Heads, S.W. 2008. The first fossil Proscopiidae with comments on the historical biogeography and evolution of the family. Palaeontology 51: 499–507.
  • Heads, S.W. 2006 . A new caddisfly larval case from the Lower Cretaceous Vectis Formation of the Isle of Wight, southern England. Proceedings of the Geologists’ Association 117: 307–310.
  • Menon, F., Heads, S.W. & Martill, D.M. 2005. New Palaeontinidae from the Lower Cretaceous Crato Formation of Brazil. Cretaceous Research 26: 837–844.
  • Menon, F. & Heads, S.W. 2005. New species of Palaeontinidae from the Lower Cretaceous Crato Formation of Brazil. Stuttgarter Beiträge zur Naturkunde, Serie B 357: 1–11.
  • Heads, S.W., Martill, D.M. & Loveridge, R.F. 2005. An exceptionally preserved antlion with colour pattern preservation from the Cretaceous of Brazil. Palaeontology 48: 1409–1417.