Lympho-epithelial Kazal-type related inhibitor 2 is a protein encoded by the SPINK9gene in humans. SPINK9 is a member of a gene familycluster located on chromosome 5q33.1, which includesSPINK5 and SPINK6. LEKTI-2 is an inhibitor of KLK5.
Desquamation
The outer layer of the epidermis is called the stratum corneum. In the stratum corneum terminally differentiatedcorneocytes are held together by corneodesmosomes. In order for desquamation to occur, corneodesmosomes need to be fully degraded. KLK5 and KLK7 are two serine proteases that degrade corneodesmosomes. LEKTI-2 regulates corneodesmosome degradation by inhibiting KLK5. In acral skin, where desquamation needs to be delayed, SPINK9expression is strongly upregulated. The resulting high level of LEKTI-2 delays corneodesmosome degradation, thereby allowing the epidermis to form a thick protective stratum corneum layer.