Upon finding an alfalfa leafcutting beeleaf cell containing a prepupa or pupa, the P. venustusfemale insert its ovipositorthrough the wall of the cell and sting the bee larva or pupa, thus paralyzing it. It then lays a number of eggs onto the paralized host, and goes on to searching for other leafcutting bee cells. The eggs hatch within 24-48 h, and the larvae immediately start feeding on the paralyzed host. If the temperatures are sufficiently cold the P. venustus pre-pupae will overwinter, but if the temperatures remain sufficiently warm they will pupate and metamorphose to adults. To exit a leaf cell, one P. venustus adult chews an exit hole, through which all the other P. venustus present within this cell will exit. Adults mate immediately, and the females then search for hosts to parasitize.