Paoliida

Paoliida is an extinct order of winged insects that lived in the late Paleozoic. Historically, both their systematic position and composition were controversial – for instance they had been considered as palaeodictyopterans, as basal Neoptera, or as stem-group of Pterygota – but recent studies have resolved them as the sister group of Dictyoptera.

Families and genera

After Prokop et al., Guan et al. and Nel & Poschmann :

†Anthracoptilidae Handlirsch, 1922
* ?†Adiphlebia Scudder, 1885
* †Afrocladus Nel, Garrouste & Prokop, 2015
* †Anthracoptilus Lameere, 1917
* †Carrizocladus Rasnitsyn in Rasnitsyn et al., 2004
* †Graticladus Novokshonov and Aristov, 2004
* †Homocladus Carpenter, 1966
* †Jarmilacladus Rasnitsyn and Aristov, 2004
* †Lodevocladus Prokop et al., 2015
* †Mesoptilus Lameere, 1917
* †Mycteroptila Rasnitsyn, 1977
* †Paracladus Carpenter, 1966
* †Pruvostia Bolton, 1921
* †Pseudomesoptilus Guan et al., 2016
* †Rhinomaloptila Rasnitsyn, 1977
* †Spargoptilon Kukalová, 1965
* †Strephocladus Scudder, 1885
* †Strephoneura Martynov, 1940
* †Strephoptilus Rasnitsyn and Aristov, 2013
* †Westphaloptilus Guan et al., 2016
†Blattinopsidae Bolton, 1925
* †Alienus Handlirsch, 1906
* †Avionblattinopsis Quispe et al., 2021
* †Balduriella Meunier, 1925
* †Blattinopsis Giebel, 1867
* †Glaphyrophlebia Handlirsch, 1906
* †Klebsiella Meunier, 1908
* †Microblattina Scudder, 1895
* †Protoblattiniella Meunier, 1912
* †Rhipidioptera Brongniart, 1894
* †Stephanopsis Kukalová, 1958
* †Westphaloblattinopsis Béthoux and Jarzembowski, 2010
†Paoliidae Handlirsch, 1906
* †Acropermula Aristov and Storozhenko, 2011
* †Aenigmidelia Sharov, 1961
* †Archidelia Sharov, 1961
* †Carbonidelia Nel, Garrouste & Jouault, 2023
* †Darekia Prokop et al., 2012
* †Herbstiala Schmidt, 1952
* †Holasicia Kukalová, 1958
* †Idelina Storozhenko, 1992
* †Kemperala Brauckmann, 1984
* †Khosaridelia Storozhenko, 1992
* †Kortshakolia Sharov, 1961
* †Kochopteron Brauckmann, 1984
* †Mertovia Prokop and Nel, 2007
* †Micaidelia Aristov 2004
* †Mongoloidelia Storozhenko, 1992
* †Olinka Kukalová, 1958
* ?†Paolekia Riek, 1976
* †Paolia Smith, 1871
* †Paoliola Handlirsch, 1919
* †Permotermopsis Martynov, 1937
* †Permula Handlirsch, 1919
* †Protoblattina Meunier, 1909
* †Pseudofouquea Handlirsch, 1906
* †Silesiapteron Prokop et al., 2013
* †Simplexpaolia Santos et al., 2023
* †Sojanidelia Storozhenko, 1992
* †Stenaropodites Martynov, 1928
* †Sustaia Kukalová, 1958
* †Sylvidelia Martynov, 1940
* †Zdenekia Kukalová, 1958