Paoliida


Paoliida is an extinct order of winged insects that lived in the late Paleozoic. Historically, both their systematic position and composition were controversial – for instance they had been considered as palaeodictyopterans, as basal Neoptera, or as stem-group of Pterygota – but recent studies have resolved them as the sister group of Dictyoptera.

Families and genera

After Prokop et al., Guan et al. and Nel & Poschmann :
  • †Anthracoptilidae Handlirsch, 1922
  • * ?†Adiphlebia Scudder, 1885
  • * †Afrocladus Nel, Garrouste & Prokop, 2015
  • * †Anthracoptilus Lameere, 1917
  • * †Carrizocladus Rasnitsyn in Rasnitsyn et al., 2004
  • * †Graticladus Novokshonov and Aristov, 2004
  • * †Homocladus Carpenter, 1966
  • * †Jarmilacladus Rasnitsyn and Aristov, 2004
  • * †Lodevocladus Prokop et al., 2015
  • * †Mesoptilus Lameere, 1917
  • * †Mycteroptila Rasnitsyn, 1977
  • * †Paracladus Carpenter, 1966
  • * †Pruvostia Bolton, 1921
  • * †Pseudomesoptilus Guan et al., 2016
  • * †Rhinomaloptila Rasnitsyn, 1977
  • * †Spargoptilon Kukalová, 1965
  • * †Strephocladus Scudder, 1885
  • * †Strephoneura Martynov, 1940
  • * †Strephoptilus Rasnitsyn and Aristov, 2013
  • * †Westphaloptilus Guan et al., 2016
  • †Blattinopsidae Bolton, 1925
  • * †Alienus Handlirsch, 1906
  • * †Avionblattinopsis Quispe et al., 2021
  • * †Balduriella Meunier, 1925
  • * †Blattinopsis Giebel, 1867
  • * †Glaphyrophlebia Handlirsch, 1906
  • * †Klebsiella Meunier, 1908
  • * †Microblattina Scudder, 1895
  • * †Protoblattiniella Meunier, 1912
  • * †Rhipidioptera Brongniart, 1894
  • * †Stephanopsis Kukalová, 1958
  • * †Westphaloblattinopsis Béthoux and Jarzembowski, 2010
  • †Paoliidae Handlirsch, 1906
  • * †Acropermula Aristov and Storozhenko, 2011
  • * †Aenigmidelia Sharov, 1961
  • * †Archidelia Sharov, 1961
  • * †Carbonidelia Nel, Garrouste & Jouault, 2023
  • * †Darekia Prokop et al., 2012
  • * †Herbstiala Schmidt, 1952
  • * †Holasicia Kukalová, 1958
  • * †Idelina Storozhenko, 1992
  • * †Kemperala Brauckmann, 1984
  • * †Khosaridelia Storozhenko, 1992
  • * †Kortshakolia Sharov, 1961
  • * †Kochopteron Brauckmann, 1984
  • * †Mertovia Prokop and Nel, 2007
  • * †Micaidelia Aristov 2004
  • * †Mongoloidelia Storozhenko, 1992
  • * †Olinka Kukalová, 1958
  • * ?†Paolekia Riek, 1976
  • * †Paolia Smith, 1871
  • * †Paoliola Handlirsch, 1919
  • * †Permotermopsis Martynov, 1937
  • * †Permula Handlirsch, 1919
  • * †Protoblattina Meunier, 1909
  • * †Pseudofouquea Handlirsch, 1906
  • * †Silesiapteron Prokop et al., 2013
  • * †Simplexpaolia Santos et al., 2023
  • * †Sojanidelia Storozhenko, 1992
  • * †Stenaropodites Martynov, 1928
  • * †Sustaia Kukalová, 1958
  • * †Sylvidelia Martynov, 1940
  • * †Zdenekia Kukalová, 1958