Nomorhamphus
Nomorhamphus is a southeast Asian genus of viviparous halfbeaks from streams, rivers and lakes in Sulawesi and the Philippines. They are all viviparous, producing small clutches of around a dozen fry about long at birth. Females are generally larger than the males. In the largest species, such as Nomorhamphus liemi, the females typically reach about in length, whereas the males reach about in length. Males are also more brightly coloured than the females. Compared with many other halfbeaks, the lower mandible, or beak, is relatively short, on females in particular barely protruding beyond the length of the upper mandible. The males of some species have short, straight beaks, but those of others have short beaks that curve downwards forming a shape often compared to a goatee beard by aquarists. N. aenigma is unique within Nomorhamphus because of its lack of lower jaw elongation.
Nomorhamphus feed extensively on small insects, either in the form of aquatic larvae or as flying insects that have fallen onto the surface of the water. They are important predators on insects such as mosquitoes, so play a role in controlling malaria. Nomorhamphus are too small to be of value as food, but they do have some value as aquarium fish.
Reproduction
Nomorhamphus species are livebearing fish that practise internal fertilisation. The male is equipped with a gonopodium-like anal fin known as a gonopodium that delivers sperm into the female. The gestation period is about six weeks. The exact mode of reproduction ranges from ovoviviparity through to viviparity, and in some species, oophagy is known, as well. Only around ten to twenty embryos are developed at a time, but at birth these are fairly large and well developed, able to take small prey, such as Daphnia, immediately after birth.Species
There are currently 22 recognized species in this genus:Nomorhamphus aenigma Kobayashi, Masengi & Yamahira, 2020- Nomorhamphus bakeri Fowler & Bean, 1922Nomorhamphus brembachi Vogt, 1978Nomorhamphus celebensis Weber & de Beaufort, 1922 Nomorhamphus ebrardtii Popta, 1912Nomorhamphus hageni Popta, 1912Nomorhamphus kolonodalensis Meisner & Louie, 2000Nomorhamphus lanceolatus Huylebrouck, Hadiaty & Herder, 2014 Nomorhamphus liemi Vogt, 1978Nomorhamphus manifesta Meisner, 2001Nomorhamphus megarrhamphus Brembach, 1982Nomorhamphus pectoralis Fowler, 1934Nomorhamphus philippina Ladiges, 1972Nomorhamphus pinnimaculatus Meisner, 2001Nomorhamphus ravnaki Brembach, 1991Nomorhamphus rex Huylebrouck, Hadiaty & Herder, 2012 Nomorhamphus rossi Meisner, 2001Nomorhamphus sagittarius Huylebrouck, Hadiaty & Herder, 2014 Nomorhamphus sanussii Brembach, 1991Nomorhamphus towoetii Ladiges, 1972Nomorhamphus versicolor Kraemer, Hadiaty & Herder, 2019Nomorhamphus vivipara Peters, 1865Nomorhamphus weberi Boulenger, 1897