Mydas fly


The Mydidae, or Mydas flies, are a cosmopolitan family of flies. It is a small family, with about 471 species described. They are generally large in size, including the largest known fly, Gauromydas heros. Many of the species, in addition to their large size, are mimics of stinging hymenopterans, especially wasps.
Most mydids are found in arid and semiarid regions of the world, but they are also found in other habitats. Mydidae are most diverse in southern Africa, but many species have also been described from North America, Chile, and Australia.

Biology

Little is known about their biology, though Zikan reported the larvae of Gauromydas heros live in the subterranean detritus "pans" of Atta ants in southern Brazil, where they appear to be feeding on detritivorous Dynastinae larvae. In the U.S., Mydas brunneus, Mydas clavatus, and Mydas tibialis larvae are predatory on deadwood-feeding scarab beetle larvae and can be found in standing and downed trees with extensive heart rot. Others are subterranean and feed on "white grubs" that attack the roots of grasses and could be potential biocontrol agents of white grubs in sod production areas. Larvae typically take two or three years to mature.
Adults of several species are avid flower visitors and act as pollinating agents. Rattlesnake master is a favorite nectar source in the Midwest. They are infrequently encountered as the adult lifespan can be quite short.

Description

Mydids are medium-sized to very large flies. The abdomen is long and cylindrical in section. It is slightly tapered apically in the male, and usually widest at segment four in the female. The second segment of the antenna forms a club. Mydids are sparsely pilose, and lack bristles except on the legs. The hind leg is much longer and stronger than both the middle leg and the fore leg and the hind femur is usually swollen and bears ventral spines. The hind tibia has an apical spur or bristles. The wings are long, and narrow to wide. Most of the veins end in the upper margin before the apex.

Classification

The classification of the family has changed fairly recently, in 1996, with the inclusion of a few genera that were previously placed in the family Apioceridae. Apiocerids have long been given the common name "flower-loving flies", but the only group of "flower-loving flies" that actually visit flowers were transferred to the Mydidae. Among the genera transferred is Rhaphiomidas, which includes one of the few Diptera on the United States' Endangered Species List, the Delhi Sands flower-loving fly.

Evolutionary history

The oldest known member of the family is Cretomydas from the Early Cretaceous Crato Formation of Brazil. The genus appears to have close affinites to the subfamily Diochlistinae, suggesting a Cretaceous diversification for the family.

Species lists

*
*
*
*
*

Genera

Afroleptomydas Bequaert, 1961Afromydas Bequaert, 1961Agaperemius Hesse, 1969Anomalomydas Papavero & Wilcox, 1974Apiophora Philippi, 1865Arenomydas Hesse, 1969 c gBaliomydas Papavero & Pimentel, 1989Belemiana Pontia, 2020Cacatuopyga Papavero & Wilcox, 1974Cephalocera c gCephalocerodes Hesse, 1969 c gCeriomydas Williston, 1898Charimydas c gChrysomydas Papavero & Pimentel, 1989Diochlistus Gerstaecker, 1868 c gDolichogaster Macquart, 1848Ectyphus Gerstaecker, 1868 c gEremohaplomydas c gEremomidas c gEumydas c gGauromydas Papavero & Pimentel, 1989Halterorchis Bezzi, 1924Haplomydas c gHessemydas Carr & Irwin, 2005Heteroleptomydas c gHeteromydas Hardy, 1944Hispanomydas c gLachnocorynus Hesse, 1969 c gLeptomydas Gerstaecker, 1868 c gMahafalymydas c gMapinguari Papavero & Wilcox, 1974Megascelus c gMessiasia d'Andretta, 1951 i c g bMidacritus c gMiltinus Gerstaecker, 1868 c gMimadelphus Hesse, 1972Mitrodetus Gerstaecker, 1868 c gMydas Fabricius, 1794Mydaselpis c gNamadytes Hesse, 1969 c gNamibimydas Hesse, 1972Nemomydas Curran, 1934 i c g bNeolaparopsis Hesse, 1969 c gNeorhaphiomidas c gNomoneura c gNomoneuroides Hesse, 1969 c gNothomydas Hesse, 1969Notosyllegomydas Hesse, 1969 c gOpomydas Curran, 1934Oreomydas Hesse, 1969 c gParamydas c gParectyphus c gPerissocerus Gerstaecker, 1868 c gPhyllomydas Bigot, 1880Plyomydas Papavero, 1971Protomydas Papavero & Pimentel, 1989Pseudonomoneura Bequaert, 1961Pseudorhopalia Wilcox & Papavero, 1971Rhaphiomidas Osten Sacken, 1877Rhopalia c gRhopaliana c gStratiomydas Papavero & Pimentel, 1989Syllegomydas c gTongamya c gVespiodes Hesse, 1969 c g
Data sources: i = ITIS, c = Catalogue of Life, g = GBIF, b = Bugguide.net