Legionaris
Legionaris is an extinct genus of ticks in the family Nuttalliellidae. The genus is currently monotypic, being represented only by the fossil species Legionaris robustus.
Systematics
Legionaris was described alongside a growing number of basal tick fossil species. The genus name, Legionaris, refers to the "ever-expanding list of genera and species that belong to the larger Nuttalliellidae family". The description of the genus was a compelling factor in collapsing other similar fossil species, including the Deinocrotonidae, into the Nuttalliellidae. Legionaris is now one of three genera placed in the Nuttalliellidae, alongside Nuttalliella and Deinocroton. The shared characters circumscribing the family are:"a pseudo-scutum/pseudo-conscutum, convoluted structure of the integument, sub-terminal mouthparts with arising of segments II–IV from the anterior segment and leg joints that extend deep into the preceding cavity, either as ball-and-socket or approximating a socket joint ".As such, the Nuttalliellidae are distinguished from the Khimairidae, which are most notably characterised by an integumented body together with a true scutum. Cladogram after Chitimia‐Dobler et al. 2022 and 2024