The holotype, now reposited at the Field Museum, was previously described by Godfrey as an aïstopod, Phlegethontia longissama. Redescription of the specimen led to its identification as a new lysorophian taxon. Infernovenator is represented only by the holotype, a nearly complete skeleton. The genus name is given for the Latin infernum to refer to the fossorial habitats of the taxon and venator. The species namehonors paleontologist Margaret Clair Steen Brough.
Anatomy
Infernovenator is diagnosed by a unique combination of features: 61 presacral vertebrae; a triangular postfrontal that contacts the tabular; a circumorbital series formed by the prefrontal, the postfrontal, the lacrimal, and the maxilla; and an ossified septomaxilla. Features shared with the lysorophian Brachydectes and not with other recumbirostrans include: postorbitalabsent and cheek emargination present; bar-like tabular-squamosal complex; and short and robust dentary.