Hakea
Hakea is a genus of about 150 species of flowering plants in the family Proteaceae, endemic to Australia. They are shrubs or small trees with leaves that are sometimes flat, otherwise circular in cross section in which case they are sometimes divided. The flowers are usually arranged in groups in leaf axils and resemble those of other genera, especially Grevillea. Hakeas have woody fruit which distinguishes them from grevilleas which have non-woody fruit which release the seeds as they mature. Hakeas are found in every state of Australia with the highest species diversity being found in the south west of Western Australia.
Description
Plants in the genus Hakea are shrubs or small trees. Some species have flat leaves, whilst others have leaves which are needle-like, in which case they are sometimes divided and sometimes have a groove on the lower surface. The flowers are arranged in groups in leaf axils and are surrounded by bracts when in bud. The flowers have both male and female parts and are borne on a short stalk called a pedicel. The sepals and petals, jointly called tepals, form a curved tube which sometimes splits open as the flower develops. The style is longer than the tepal tube and is curved before its tip is released. When released, the tip of the style is a pollen-presenter. The fruit of hakeas is woody and persists on the plant until burned in a bushfire or until the plant dies. The fruit then splits open to release two winged seeds.Hakeas are similar to other plants in the Family Proteaceae, but have undivided leaves arranged alternately, sessile flowers arranged in loose groups in the axils of leaves or bracts, unlike those in the Banksia. Hakeas are similar to species of Grevillea but are distinguished from them in having persistent, woody fruits., and the ovary and style are glabrous.
Taxonomy and naming
The genus Hakea was first formally described in 1797 by Heinrich Schrader and Johann Christoph Wendland and the description was published in Sertum Hannoveranum. The genus is named after Baron Christian Ludwig von Hake, an 18th-century German patron of botany.Distribution
Hakea are endemic to Australia, where they can be found in all six states. Some species, such as Hakea sericea, have been introduced elsewhere, where they have become naturalized and may be considered invasive. Introduced Hakea populations occur in New Zealand, Norfolk Island, South Africa and Portugal.Horticulture
Hakeas are popular ornamental plants in gardens in Australia, and in many locations are as common as grevilleas and banksias. Several hybrids and cultivars have been developed, including Hakea 'Burrendong Beauty'. They are best grown in beds of light soil which are watered but still well drained.Some showy western species, such as Hakea multilineata, H. francisiana and H. bucculenta, require grafting onto hardy stock such as Hakea salicifolia for growing in more humid climates, as they are sensitive to dieback.
Many species, particularly eastern Australian species, are notable for their hardiness, to the point they have become weedy. Hakea gibbosa, H. sericea, and H. drupacea have been weeds in South Africa, Hakea laurina has become naturalized in the eastern states of Australia and is considered an environmental weed, and H. salicifolia, H. gibbosa, and H. sericea are invasive weeds in New Zealand.
List of species
The following is a list of Hakea species recognised by the Australian Plant Census, except for Hakea asperma which is recognised by the Royal Botanic Gardens Victoria:Hakea actites W.R.Barker – mulloway needlebush, wallum hakeaHakea aculeata A.S.George – column hakeaHakea acuminata HaegiHakea adnata R.Br.Hakea aenigma W.R.Barker & Haegi – enigma hakeaHakea ambigua Meisn.Hakea amplexicaulis R.Br. – prickly hakeaHakea anadenia HaegiHakea arborescens R.Br. – common hakea, yellow hakeaHakea archaeoides W.R.Barker- Hakea asperma Molyneux & Forrester – native dog hakeaHakea auriculata Meisn.Hakea bakeriana F.Muell. & MaidenHakea baxteri R.Br. – fan hakeaHakea bicornata R.M.BarkerHakea brachyptera Meisn. – short-winged hakeaHakea brownii Meisn. – fan-leaf hakeaHakea bucculenta C.A.Gardner – red pokersHakea candolleana Meisn.Hakea carinata F.Muell. ex Meisn.Hakea ceratophylla R.Br. – horned-leaf hakea, staghorn hakeaHakea chordophylla F.Muell. – bootlace oak, bootlace tree, corkwood, bull oakHakea chromatropa A.S.George & R.M.BarkerHakea cinerea R.Br.Hakea circumalata Meisn.Hakea clavata Labill. – coastal hakeaHakea collina C.T.WhiteHakea commutata F.Muell.Hakea conchifolia Hook.f. – shell-leaved hakeaHakea constablei L.A.S.JohnsonHakea corymbosa R.Br. – cauliflower hakeaHakea costata Meisn. – ribbed hakeaHakea cristata R.Br.Hakea cucullata R.Br. – hood-leaved hakea, scallop hakeaHakea cyclocarpa Lindl. – ramshorn hakeaHakea cycloptera R.Br.Hakea cygnus Lamont
- * Hakea cygnus Lamont subsp. cygnus – swan-fruit hakea
- * Hakea cygna subsp. needlei LamontHakea dactyloides Gaertn. – finger hakeaHakea decurrens R.Br.
- * Hakea decurrens R.Br. subsp. decurrens
- * Hakea decurrens subsp. physocarpa W.R.Barker
- * Hakea decurrens subsp. platytaenia W.R.BarkerHakea denticulata R.Br. – stinking RogerHakea divaricata L.A.S.Johnson – needlewood, corkbark tree, fork-leaved corkwoodHakea dohertyi HaegiHakea drupacea Roem. & Schult.Hakea ednieana Tate – Flinders Range hakea, yandenaHakea elliptica R.Br. – oval-leaf hakea Hakea eneabba HaegiHakea epiglottis Labill.
- * Hakea epiglottis Labill. subsp. epiglottis
- * Hakea epiglottis subsp. milliganii R.M.BarkerHakea erecta LamontHakea eriantha R.Br. – tree hakeaHakea erinacea Meisn. – hedgehog hakea, porcupine hakeaHakea eyreana McGill – straggly corkbarkHakea falcata R.Br. – sickle hakeaHakea ferruginea SweetHakea flabellifolia Meisn. – fan-leaved hakea, wedge hakeaHakea florida R.Br.Hakea florulenta Meisn.Hakea francisiana F.Muell. – emu tree, grass-leaf hakea, bottlebrush hakeaHakea fraseri R.Br. – corkwood oakHakea gibbosa Cav. – hairy hakea, rock hakeaHakea gilbertii KippistHakea grammatophylla F.Muell.Hakea hastata HaegiHakea hookeriana Meisn. – Barren Range hakeaHakea horrida R.M.BarkerHakea ilicifolia R.Br.Hakea incrassata R.Br. – marble hakeaHakea invaginata B.L.BurttHakea ivoryi F.M.Bailey – Ivory's hakea, corkwood, corkbark treeHakea kippistiana Meisn.Hakea laevipes Gand.
- * Hakea laevipes subsp. graniticola Haegi
- * Hakea laevipes Gand. subsp. laevipes Hakea lasiantha R.Br. – woolly-flowered hakeaHakea lasianthoides RyeHakea lasiocarpha R.Br. – long-styled hakeaHakea laurina R.Br. – kodjet, pin-cushion hakea, emu bushHakea lehmanniana Meisn. – blue hakeaHakea leucoptera R.Br. – silver needlewood, needle hakea, pin bush, water tree, booldoobah
- * Hakea leucoptera R.Br. subsp. leucoptera
- * Hakea leucoptera subsp. sericipes W.R.BarkerHakea linearis R.Br.Hakea lissocarpha R.Br. – honey bush, honeybush hakeaHakea lissosperma R.Br. – needle bush, mountain needlewoodHakea longiflora R.M.BarkerHakea loranthifolia Meisn.Hakea lorea R.Br. – bootlace oak, cork tree
- * Hakea lorea subsp. borealis W.R.Barker
- * Hakea lorea R.Br. subsp. loreaHakea maconochieana HaegiHakea macraeana F.Muell. – willow needlewood, Macrae's hakeaHakea macrocarpa A.Cunn. ex R.Br. – dogwood hakeaHakea macrorrhyncha W.R.BarkerHakea marginata R.Br.Hakea megadenia R.M.BarkerHakea megalosperma Meisn. – Lesueur hakeaHakea meisneriana KippistHakea microcarpa R.Br. – small-fruit hakeaHakea minyma MaconochieHakea mitchellii Meisn.Hakea multilineata Meisn. – grass-leaf hakeaHakea myrtoides Meisn. – myrtle hakeaHakea neospathulata Hakea neurophylla Meisn. – pink-flowered hakeaHakea newbeyana R.M.BarkerHakea nitida R.Br. – frog hakea, shining hakeaHakea nodosa R.Br. – yellow hakeaHakea obliqua R.Br. – needles and corks
- * Hakea obliqua R.Br. subsp. obliqua
- * Hakea obliqua subsp. parviflora R.M.BarkerHakea obtusa Meisn.Hakea ochroptera W.R.BarkerHakea oldfieldii Benth.Hakea oleifolia R.Br. – dungyn, olive-leaf hakeaHakea oligoneura K.A.Sheph. & R.M.BarkerHakea orthorrhyncha F.Muell. – bird beak hakea
- * Hakea orthorrhyncha var. filiformis F.Muell. ex Benth.
- * Hakea orthorrhyncha F.Muell. ex Benth. var. orthorrhynchaHakea pachyphylla Sieber ex Spreng.Hakea pandanicarpa R.Br.
- * Hakea pandanicarpa subsp. crassifolia R.M.Barker
- * Hakea pandanicarpaR.Br. subsp. pandanicarpaHakea pedunculata F.Muell.Hakea pendens R.M.BarkerHakea persiehana F.Muell.Hakea petiolaris Meisn. – sea-urchin hakea
- * Hakea petiolaris subsp. angusta Haegi
- * Hakea petiolaris Meisn. subsp. petiolaris
- * Hakea petiolaris subsp. trichophylla HaegiHakea platysperma Hook. – cricket ball hakea, woody peachHakea polyanthema DielsHakea preissii Meisn. – needle tree, needle bush, Christmas hakeaHakea pritzelii DielsHakea propinqua A.Cunn.Hakea prostrata R.Br. – harsh hakeaHakea psilorrhyncha R.M.BarkerHakea pulvinifera L.A.S.Johnson - Lake Keepit hakea Hakea purpurea Hook.Hakea pycnoneura Meisn.Hakea recurva Meisn. – djarnokmurd
- * Hakea recurva subsp. arida W.R.Barker & R.M.Barker
- * Hakea recurva Meisn. subsp. recurvaHakea repullulans H.M.LeeHakea rhombales F.Muell. – walukaraHakea rigida C.A.Gardner ex HaegiHakea rostrata F.Muell. ex Meisn.Hakea rugosa R.Br. – wrinkled hakeaHakea ruscifolia Labill. – candle hakeaHakea salicifolia B.L.Burtt – willow-leaved hakea
- * Hakea salicifolia subsp. angustifolia W.R.Barker
- * Hakea salicifolia B.L.Burtt subsp. salicifoliaHakea scoparia Meisn.
- * Hakea scoparia Meisn. subsp. scoparia
- * Hkea scoparia subsp. trycherica HaegiHakea sericea Schrad. & J.C.Wendl. – needlebush, silky hakeaHakea smilacifolia Meisn.Hakea standleyensis MaconochieHakea stenocarpa R.Br.Hakea stenophylla A.Cunn. ex R.Br.
- * Hakea stenophylla subsp. notialis R.M.Barker
- * Hakea stenophylla A.Cunn. ex R.Br. subsp. stenophyllaHakea strumosa Meisn.Hakea subsulcata Meisn.Hakea sulcata R.Br.Hakea tephrosperma R.Br. – hooked needlewoodHakea teretifolia Britten – dagger hakeaHakea teretifolia subsp. hirsuta R.M.BarkerHakea teretifolia Britten subsp. teretifoliaHakea trifurcata R.Br. – two-leaf, two-leaved hakea, kerosine bushHakea trineura F.Muell. F.Muell.Hakea tuberculata R.Br.Hakea ulicina R.Br. - furze hakea Hakea undulata R.Br. – wavy-leaved hakeaHakea varia R.Br. –variable-leaved hakeaHakea verrucosa F.Muell.Hakea victoria J.Drumm. – lantern hakea, royal hakeaHakea vittata R.Br.