Euphorinae


The Euphorinae are a large subfamily of Braconidae parasitoid wasps. Some species have been used for biological pest control. They are sister group to the Meteorinae.
There are over 1270 species of Euphorinae.

Description and distribution

Euphorines are small, usually dark colored wasps. They are non-cyclostomes. Euphorines are found worldwide.

Biology

Euphorines are solitary or rarely gregarious koinobiont endoparasitoids. Unlike most other parasitoid wasps, Euphorinae have a broad host range and attack adult insects or nymphs of hemimetabolous insects.
Wasps of the tribe Dinocampini parasitize adult beetles.

Taxonomy and phylogeny

Shaw divided Euphorinae into three tribes, Cosmophorini and Meteorini with one genus each and Euphorini containing 33 genera. In 1997, Shaw proposed 9 tribes and 31 genera of Euphorinae. By Yu et al., the list of representative tribes of Euphorinae had grown to 16: Centistini, Cosmophorini, Cryptoxilonini, Dinocampini, Euphorini, Helorimorphini, Mannokeraiini, Meteorini, Myiocephalini, Neoneurini, †Oncometeorini, Perilitini, Planitorini, Proclithrophorini, Syntretini, and Tainitermini. Mannokeraiini was synonymised under Planitorini by van Achterberg et al.. Stigenberg et al. treated 52 genera in 14 extant tribes, elevating Pygostolini from a subtribe of Centistini, synonymizing Cryptoxilonini under Cormophorini, synonymizing Proclithrophorini under Townesilitini, and removing Tainitermini as not nested within Euphorinae. Chen & van Achterberg included the additional tribe of Eadyini and removed Proclithrophorini from synonymy. In 2021, Stigenberg & van Achterberg returned Proclithrophorini to synonymy under Townesilitini. Bendixen & Shaw elevated the Meteorini again to subfamily status as Meteorinae rather than treat them as basal Euphorinae.
The present Euphorinae thus contains 14 extant tribes: Centistini, Cosmophorini, Dinocampini, Eadyini, Ecnomiini, Euphorini, Helorimorphini, Myiocephalini, Neoneurini, Perilitini, Planitorini, Pygostolini, Syntretini, and Townesilitini.
Belokobylskij recognize two extinct tribes, Oncometeorini and Prosyntretini.

Genera

The following 56 extant genera and 6 extinct genera belong to the subfamily Euphorinae:Stenope van Achterberg & Reshchikov, 2018

Tribe [Centistini] Čapek, 1970

Allurus Förster, 1862Asiacentistes Belokobylskij, 1995Centistes Haliday, 1835Centistoides van Achterberg, 1992
  • Parasyrrhizus Brues, 1933

Tribe [Cosmophorini] Čapek, 1958

Cosmophorus Ratzeburg, 1848Cryptoxilos Viereck, 1911Plynops Shaw, 1996Ropalophorus Curtis, 1837Sinuatophorus van Achterberg, 2000Tuberidelus Chen & van Achterberg, 1997

Tribe [Dinocampini] Shaw, 1985

Betelgeuse Shaw, 1988Centistina Enderlein, 1912Dinocampus Förster, 1862Ecclitura Kokujev, 1902Yanayacu Zhang & Chen, 2015

Tribe [Eadyini] van Achterberg, 2000

Eadya Huddleston & Short, 1978

Tribe [Ecnomiini] van Achterberg, 1985

Ecnomios Mason, 1979Korecnomios Park & van Achterberg, 1994

Tribe [Euphorini] Shaw, 1985

Leiophron Nees von Esenbeck, 1818Mama Belokobylskij, 2000Peristenus Förster, 1862

Tribe [Helorimorphini] Schmiedeknecht, 1907

Aridelus Marshall, 1887Chrysopophthorus Goidanich, 1948Holdawayella Loan 1967Wesmaelia Förster, 1862

Tribe [Myiocephalini] Chen & van Achterberg, 1997

Myiocephalus Marshall, 1897

Tribe [Neoneurini] Bengtsson, 1918

Elasmosoma Ruthe, 1858
  • Elasmosomites Brues, 1933Euneoneurus Tobias & Yuldashev, 1979Kollasmosoma van Achterberg & Argaman, 1993Neoneurus Haliday, 1838Parelasmosoma Tobias & Yuldashev, 1979

†Tribe [Oncometeorini] Tobias, 1987

  • Oncometeorus Tobias, 1987

Tribe [Perilitini] Förster, 1862

  • Meteorites Brues, 1939Microctonus Wesmael, 1835Notioperilitus Belokobylskij, 2018
  • Onychoura Brues, 1933Orionis Shaw, 1987Perilitus Nees, 1818Rilipertus Haeselbarth, 1996Spathicopis van Achterberg, 1977Stenothremma Shaw, 1984

Tribe [Planitorini] van Achterberg, 1995

Mannokeraia van Achterberg, 1995Paramannokeraia van Achterberg & Quicke, 2017Planitorus van Achterberg, 1995

†Tribe [Prosyntretini] Tobias, 1987

  • Prosyntretus Tobias, 1987

Tribe [Pygostolini] Belokobylskij, 2000

Litostolus van Achterberg, 1985Pygostolus Haliday, 1833

Tribe [Syntretini] Shaw, 1985

Bracteodes De Saeger, 1946Sculptosyntretus Belokobylskij, 1993Syntretellus De Saeger, 1946Syntretomorpha Papp, 1962Syntretoriana Parrott, 1953Syntretus Förster, 1862

Tribe [Townesilitini] Shaw, 1985

Gretiella Stigenberg, 2021Heia Chen & van Achterberg, 1997Marshiella Shaw, 1985Proclithrophorus Tobias & Belokobylskij, 1981Streblocera Westwood, 1833Townesilitus Haeselbarth & Loan, 1983