Emberiza


The buntings are a group of Old World passerine birds forming the genus Emberiza, the only genus in the family Emberizidae. The family contains 44 species. They are seed-eating birds with stubby, conical bills.

Taxonomy

The family Emberizidae was formerly much larger and included the species now placed in the Passerellidae and Calcariidae. Molecular phylogenetic studies found that the large family consisted of distinct clades that were better treated as separate families.
The genus Emberiza is now the only genus placed in the family Emberizidae. The genus was introduced by the Swedish naturalist Carl Linnaeus in 1758 in the tenth edition of his Systema Naturae. The type species was subsequently designated as the yellowhammer. The genus name Emberiza is from Old German Embritz, a bunting. The origin of the English "bunting" is unknown.
A 2008 genetic study found that three emberizid species that were placed in their own monotypic genera clustered within the Emberiza. These were the crested bunting, the slaty bunting, and the corn bunting. All three species are now included in the genus Emberiza.
A large DNA-based study of the passerines published in 2019 found that the buntings are most closely related to the longspurs and snow buntings in the family Calcariidae.
Ornithologists Edward Dickinson and Leslie Christidis in the fourth edition of the Howard and Moore Complete Checklist of the Birds of the World chose to split up Emberiza and recognise the genera Fringillaria, Melophus, Granativora, Emberiza, and Schoeniclus. Their example has not been followed by the online version of the Handbook of the Birds of the World nor by Frank Gill and David Donsker in the list of world birds that they maintain on behalf of the International Ornithologists' Union. The British Ornithologists' Union has argued that splitting the genus provides little benefit and destabilizes the nomenclature.
Species in the New World genus Passerina include the word "bunting" in their common names, but are now classed in the family Cardinalidae.
The family is divided into four major clades. The species in Clade I are mainly African while those in Clades II to IV are Palearctic:
The cladogram below is based on a study published in 2021. The phylogenetic relationships of two African species, the brown-rumped bunting and Vincent's bunting, were not determined.

List of species

The genus contains 44 species.
ImageCommon nameScientific nameDistribution
Crested buntingEmberiza lathamiSoutheast Asia.
Slaty buntingEmberiza siemsseniChina.
Corn buntingEmberiza calandraWestern Europe and North Africa across to northwestern China.
YellowhammerEmberiza citrinellasoutheast England and most of Europe east to the northwestern corner of Russia and western Ukraine.
Pine buntingEmberiza leucocephalosAsia
Rock buntingEmberiza cianorthwest Africa, southern Europe east to central Asia, and the Himalayas
Godlewski's buntingEmberiza godlewskiiChina, Pakistan, India, Kazakhstan, Mongolia, Myanmar, and Russia.
Meadow buntingEmberiza cioidessouthern Siberia, northern and eastern China, eastern Kazakhstan, Kyrgyzstan, Mongolia, Korea and Japan.
White-capped buntingEmberiza stewartiAfghanistan, India, Iran, Kazakhstan, Kyrgyzstan, Nepal, Pakistan, Tajikistan, Turkmenistan, and Uzbekistan.
Jankowski's buntingEmberiza jankowskiiRussian Far East, Manchuria and far northeastern Korea
Grey-necked buntingEmberiza buchananiCaspian Sea to the Altai Mountains in Central Asia
Cinereous buntingEmberiza cineraceasouthern Turkey and southern Iran
Ortolan buntingEmberiza hortulanaEuropean countries and western Asia
Cretzschmar's buntingEmberiza caesiaGreece, Turkey, Cyprus and the Levant.
Cirl buntingEmberiza cirlussouthern Europe, on the Mediterranean islands and in north Africa
Striolated buntingEmberiza striolataChad, east through south-west Asia to north-western India, Africa
House buntingEmberiza saharinorthwestern Africa from Morocco south to Mali and east to Chad.
Lark-like buntingEmberiza impetuaniAngola, Botswana, Lesotho, Namibia, South Africa, and Zimbabwe.
Cinnamon-breasted buntingEmberiza tahapisimainland sub-Saharan Africa
Gosling's buntingEmberiza goslingiMauritania and Senegal to south-western Sudan and north-eastern Democratic Republic of Congo.
Socotra buntingEmberiza socotranaYemen
Cape buntingEmberiza capensissouthern Africa from south-western Angola, eastern Zambia, Zimbabwe and southern Tanzania to the Cape.
Tristram's buntingEmberiza tristramieastern Manchuria and the Russian Far East and winters in central and southern China.
Chestnut-eared buntingEmberiza fucatathe Himalayas locally across China to south-eastern Siberia, Korea and northern Japan
Little buntingEmberiza pusillanorth-east of Europe and northern Eurosiberia to the Russian Far East and northern India, southern China and the northern parts of south-east Asia.
Yellow-browed buntingEmberiza chrysophryseastern Siberia, China
Rustic buntingEmberiza rusticasouth-east Asia, Japan, Korea, and eastern China.
Yellow-throated buntingEmberiza elegansChina, Japan, Korea, Myanmar, Russia, and Taiwan.
Yellow-breasted buntingEmberiza aureolaFinland to Bering Sea migrating to Indochina
Somali buntingEmberiza poliopleuraEthiopia, Kenya, Somalia, South Sudan, Tanzania, and Uganda
Golden-breasted buntingEmberiza flaviventrisAfrica south of the Sahara
Brown-rumped buntingEmberiza affinisSenegal to Sudan and Uganda
Cabanis's buntingEmberiza cabanisisub-Saharan Africa
Chestnut buntingEmberiza rutilaSiberia, northern Mongolia and north-eastern China.
Tibetan buntingEmberiza koslowiTibet
Black-headed buntingEmberiza melanocephalaJapan, China, Hong Kong, Thailand, Laos, South Korea and Malaysia
Red-headed buntingEmberiza brunicepsAsia-Afghanistan, Iran, Kazakhstan, Kyrgyzstan, Mongolia; Russian Federation, Tajikistan, Turkmenistan, Uzbekistan.
Yellow buntingEmberiza sulphurataJapan
Black-faced buntingEmberiza spodocephalasouthern Siberia across to northern China.
Masked buntingEmberiza personataSakhalin, the Kuril Islands, and Japan.
Grey buntingEmberiza variabilisKamchatka, Sakhalin, Kuril Islands and northern Japan
Pallas's reed buntingEmberiza pallasinortheast European Russia, north Kamchatka
Ochre-rumped buntingEmberiza yessoensiseastern Mongolia, northeast China and Ussuriland
Common reed buntingEmberiza schoeniclusEurope

Extinct species have been described:
  • Long-legged bunting
  • Emberiza shaamarica
  • Emberiza polgardiensis
  • Emberiza media
  • Emberiza parva
  • Emberiza gaspariskii
  • Emberiza bartoki
Emberiza pannonica from the upper Miocene of Hungary is also referred to this genus, but was later found to be a member of Muscicapidae.