Emarginula


Emarginula is a genus of small keyhole limpets, marine gastropod molluscs in the family Fissurellidae.

Description

The oval shell is obliquely conical with a backward-pointing recurved apex. There is a deep incision in the anterior margin. A distinct anal fasciole extends upwards from this incision. The surface is latticed, sculptured with radial ribs. There is no septum inside.

Radula

The radula has a dental formula of .1.4.1.4.1., with a wide, rectangular, acuspate rachidian tooth.
The various genera have a few dozen rows of teeth.

Species

According to the World Register of Marine Species, species within the genus Emarginula include:
  • Emarginula adamsiana Sowerby in Adams & Sowerby, 1863
  • Emarginula adriatica Costa O.G., 1829
  • Emarginula agulhasensis Thiele, 1925
  • Emarginula alba Watson, 1897
  • Emarginula amyda Shikama, 1962
  • Emarginula angusta McLean, 1970
  • Emarginula annielangleitae Poppe & Tagaro, 2020
  • Emarginula apolonia Pacaud, 2015
  • † Emarginula auversiensis Deshayes, 1861
  • Emarginula bajula Hedley, 1913
  • Emarginula balicasagensis Poppe & Tagaro, 2020
  • Emarginula bellula A. Adams, 1852
  • Emarginula bicancellata Montrouzier, 1860
  • Emarginula bonfittoi Smriglio & Mariottini, 2001
  • Emarginula boucheti Poppe & Tagaro, 2020
  • † Emarginula brebioni Ceulemans, Van Dingenen & Landau, 2016
  • Emarginula camilla Melvill & Standen, 1903
  • Emarginula candida Adams, 1852
  • Emarginula capuloidea G. Nevill & H. Nevill, 1869
  • † Emarginula carezi de Laubrière, 1881
  • Emarginula choristes Dall, 1925
  • Emarginula christiaensi Piani, 1985
  • Emarginula circumalbum Poppe & Tagaro, 2020
  • † Emarginula clathrata Deshayes, 1824
  • † Emarginula clathrataeformis Eichwald, 1830
  • Emarginula colipanoae Poppe & Tagaro, 2020
  • Emarginula compta Habe, 1953
  • Emarginula concinna Adams, 1852
  • Emarginula connectens Thiele, 1915
  • Emarginula connelli Kilburn, 1978
  • Emarginula convexa Hedley, 1907
  • † Emarginula costata Lamarck, 1803
  • Emarginula costulata Deshayes, 1863
  • Emarginula crassa Sowerby, 1812
  • Emarginula crassicostata Sowerby in Adams & Sowerby, 1863
  • Emarginula cucullata A. Adams, 1852
  • Emarginula curvamen Iredale, 1925
  • Emarginula curvata Schepman, 1908
  • Emarginula cuvieri Audouin, 1827
  • Emarginula decorata Deshayes, 1863
  • † Emarginula delicatissima Chapman & Gabriel, 1923
  • Emarginula delonguevilleae Poppe & Tagaro, 2020
  • † Emarginula dennanti Chapman & Gabriel, 1923
  • Emarginula devota Thiele, 1915
  • Emarginula dictya McLean, 1970
  • † Emarginula didactica Ludbrook, 1956
  • † Emarginula dilatoria Ludbrook, 1956
  • Emarginula dilecta Adams, 1852
  • Emarginula divae
  • Emarginula dubia Schepman, 1908
  • Emarginula dujardini Dollfus & Dautzenberg, 1886 †
  • † Emarginula espibosensis Lozouet, 1999
  • † Emarginula exigua Trautschold, 1866
  • Emarginula fenestrella Deshayes, 1863
  • Emarginula fissura - slit limpet
  • † Emarginula fossorensis Lozouet, 1999
  • Emarginula foveolata Schepman, 1908
  • * Emarginula foveolata fugitai Habe, 1953
  • † Emarginula foveolata Gerasimov, 1955
  • Emarginula fragilis Yokoyama, 1920
  • Emarginula fuliginea Adams, 1851
  • Emarginula gabensis
  • Emarginula galericulata A. Adams, 1852
  • † Emarginula galeriformis Marwick, 1928
  • † Emarginula gigantea Poppe, 2008
  • †Emarginula guerangeri A. d'Orbigny, 1843
  • Emarginula gustavi Poppe & Tagaro, 2020
  • Emarginula hataii Habe, 1953
  • Emarginula hawaiiensis Dall, 1895
  • † Emarginula haweraensis Powell, 1931
  • Emarginula hosoyai Habe, 1953
  • Emarginula huzardii
  • Emarginula icosisculpta Simone & Cunha, 2014
  • Emarginula imaizumi Dall, 1926
  • Emarginula imella Dall, 1926
  • Emarginula janae Cossignani, 2020
  • Emarginula japonica Sowerby in Adams & Sowerby, 1863
  • Emarginula jungcheni K.-Y. Lai, 2019
  • † Emarginula kaiparica Laws, 1939
  • † Emarginula kaiwarensis Bandel, Gründel & P. A. Maxwell, 2000 †
  • † Emarginula karuna Das, Bardhan & Lahiri, 1999
  • Emarginula kashimaensis Shikama, 1962
  • † Emarginula komitica Laws, 1939
  • Emarginula koon Kilburn, 1978
  • † Emarginula ladowae Eichman, 1955
  • Emarginula lata J.R.C. Quoy & J.P. Gaimard, 1834
  • Emarginula lentiginosa D. G. Herbert, 2024
  • Emarginula liuzzii Poppe & Tagaro, 2020
  • Emarginula longifissa Sowerby, 1866
  • Emarginula lorenzoi Giusti & Micali, 2019
  • Emarginula macclurgi Kilburn, 1978
  • Emarginula mactanensis Poppe & Tagaro, 2020
  • Emarginula maculata Adams in Adams & Sowerby, 1863
  • Emarginula margarita Poppe & Tagaro, 2020
  • † Emarginula mariae Cossmann, 1888
  • † Emarginula maudensis Chapman & Gabriel, 1923
  • Emarginula micans Adams, 1852
  • Emarginula montrouzieri Souverbie, 1872
  • Emarginula multisquamosa Schepman, 1908
  • Emarginula multistriata Jeffreys, 1882
  • Emarginula natalensis Barnard, 1963
  • Emarginula nodulicostata D. G. Herbert, 2024
  • Emarginula nordica Pérez Farfante, 1947 - northern emarginula
  • † Emarginula oblonga Sandberger, 1859
  • Emarginula obovata Adams, 1852
  • Emarginula octaviana Coen, 1939
  • Emarginula oppressa Barnard, 1963
  • Emarginula paivana
  • Emarginula papilionacea G. Nevill & H. Nevill, 1869
  • Emarginula patula Cotton, 1930
  • † Emarginula paucicostata Laws, 1936
  • Emarginula paucipunctata Schepman, 1908
  • † Emarginula pelagica Passy, 1832
  • Emarginula philippinensis Poppe & Tagaro, 2020
  • Emarginula phrixodes Dall, 1927 - ruffled emarginula
  • Emarginula pittensis Marwick, 1928
  • Emarginula poppeorum Romani & Crocetta, 2017
  • † Emarginula praesicula Lozouet, 1999
  • Emarginula puncticulata Adams, 1852
  • Emarginula punctulum Piani, 1980
  • † Emarginula punctura S. Wood, 1848
  • Emarginula pustula Thiele in Küster, 1913
  • Emarginula quadrata Poppe & Tagaro, 2020
  • † Emarginula radiocostata Kiel & Bandel, 2000
  • Emarginula regia Habe, 1953
  • Emarginula retecosa Adams, 1851
  • Emarginula retrogyra D. G. Herbert, 2024
  • Emarginula rollandi P. Fischer, 1857
  • Emarginula rosea Bell, 1824
  • Emarginula salebrosa D. G. Herbert, 2024
  • † Emarginula sanctaecaatharine Passy, 1832
  • Emarginula scabricostata A. Adams, 1852
  • Emarginula scabriuscula Adams, 1852
  • Emarginula sicula Gray, 1825 - dagger emarginula
  • Emarginula sinica Lu, 1986
  • Emarginula solidula Costa, 1829
  • Emarginula souverbiana Pilsbry, 1890
  • Emarginula spinosa Deshayes, 1863
  • † Emarginula squammata Grateloup, 1836
  • † Emarginula striatissima Traub, 1979
  • Emarginula striatula Quoy and Gaimard, 1834 - dagger emarginula
  • Emarginula sublaevis Schepman, 1908
  • Emarginula subtilitexta Verco, 1908
  • Emarginula superba Hedley, 1906
  • Emarginula suspira Simone & Cunha, 2014
  • Emarginula tenera Locard, 1892
  • † Emarginula teneraeformis Lozouet, 1999
  • Emarginula tenuicostata Sowerby, 1863
  • Emarginula teramachii Habe, 1953
  • Emarginula textilis Gould, 1859 in 1859-61
  • Emarginula thomasi Crosse, 1864
  • Emarginula thorektes Kilburn, 1978
  • Emarginula tosaensis Habe, 1953
  • †Emarginula transenna Tenison Woods, 1877
  • Emarginula tuberculosa Libassi, 1859 - tuberculate emarginula
  • Emarginula undulata Melvill & Standen, 1903
  • Emarginula velascoensis Shasky, 1961
  • Emarginula velascoi Rehder, 1980
  • Emarginula viridibrunneis Poppe & Tagaro, 2020
  • Emarginula viridicana Herbert & Kilburn, 1985
  • Emarginula viridis Poppe & Tagaro, 2020
  • † Emarginula wannonensis G. F. Harris, 1897
  • † Emarginula washingtoniana Squires & Goedert, 1994
  • Emarginula xishaensis Lu, 1986
  • Emarginula yangeorum Poppe & Tagaro, 2020

    Synonyms

  • Emarginula altilis Gould, 1859: synonym of Variemarginula punctata
  • Emarginula amitina Iredale, 1925: synonym of Emarginula dilecta A. Adams, 1852
  • Emarginula angustata Thiele, 1915: synonym of Tugali decussata A. Adams, 1852
  • Emarginula arabica A. Adams, 1852: synonym of Octomarginula arabica
  • Emarginula arconatii Issel, 1869: synonym of Montfortista panhi
  • Emarginula australis Quoy & Gaimard, 1834: synonym of Montfortia subemarginata
  • Emarginula bella Gabb, 1865: synonym of Scelidotoma bella
  • Emarginula biangulata Sowerby, 1901: synonym of Emarginella biangulata
  • † Emarginula blainvillii Defrance, 1825: synonym of † Rimula blainvillii
  • Emarginula brevirimata Deshayes, 1862: synonym of Montfortula brevirimata
  • Emarginula clathrata A. Adams & Reeve, 1850: synonym of Montfortista panhi
  • Emarginula clathrata Pease, 1863: synonym of Emarginula dilecta A. Adams, 1852
  • Emarginula conica Lamarck, 1801: synonym of Emarginula fissura
  • Emarginula conica Schumacher, 1817 non Lamarck, 1801: synonym of Emarginula rosea Bell, 1824
  • Emarginula dentigera Heilprin, 1889: synonym of Hemimarginula dentigera
  • Emarginula elata Locard, 1898: synonym of Emarginula christiaensi Piani, 1985
  • Emarginula elongata O. G. Costa, 1829: synonym of Emarginula octaviana Coen, 1939
  • Emarginula eximia A. Adams, 1852: synonym of Emarginella eximia
  • Emarginula flindersi Cotton, 1930: synonym of Emarginula convexa Hedley, 1907
  • Emarginula fuliginea Kuroda, 1941: synonym of Variemarginula variegata
  • Emarginula guernei Dautzenberg & Fischer, 1896: synonym of Emarginula tuberculosa Libassi, 1859
  • Emarginula hedleyi Thiele, 1915: synonym of Emarginula candida A. Adams, 1852
  • Emarginula incisula: synonym of Emarginella incisura
  • Emarginula incisura A. Adams, 1852: synonym of Emarginella incisura
  • Emarginula kimberti Cotton, 1930: synonym of Laeviemarginula kimberti
  • Emarginula modesta H. Adams, 1872: synonym of Hemimarginula modesta
  • Emarginula muelleri Forbes & Hanley, 1849: synonym of Emarginula fissura
  • Emarginula ostheimerae Abbott, 1958: synonym of Octomarginula ostheimerae
  • Emarginula panhi Quoy & Gaimard, 1834: synonym of Montfortista panhi
  • Emarginula papillosa Risso 1826: synonym of Emarginula huzardii Payraudeau, 1826
  • Emarginula parmophoidea Quoy & Gaimard, 1834: synonym of Tugali elegans Gray, 1843
  • Emarginula peasei Thiele, 1915: synonym of Emarginula dilecta A. Adams, 1852
  • Emarginula phrygium Herbert & Kilburn, 1985: synonym of Agariste phrygium
  • Emarginula picta Dunker, 1860: synonym of Montfortula picta
  • Emarginula pileata Gould, 1859: synonym of Variemarginula punctata
  • Emarginula pileum Heilprin, 1889: synonym of Hemimarginula pileum
  • Emarginula planulata A. Adams, 1852: synonym of Emarginella planulata
  • Emarginula polygonalis A. Adams, 1852: synonym of Hemitoma polygonalis
  • Emarginula pulchreclathrata Tomlin, 1932: synonym of Emarginula agulhasensis Thiele, 1925
  • Emarginula pumila A. Adams, 1852: synonym of Hemimarginula pumila
  • Emarginula punctata A. Adams, 1852: synonym of Variemarginula punctata
  • Emarginula reticulata Sowerby, 1813: synonym of Emarginula fissura
  • Emarginula rugosa Quoy & Gaimard, 1834: synonym of Montfortula rugosa
  • Emarginula scutellata Deshayes, 1863: synonym of Octomarginula scutellata
  • Emarginula sibogae Schepman, 1908: synonym of Emarginella sibogae
  • Emarginula simpla Christiaens, 1987: synonym of Hemimarginula simpla
  • Emarginula stellata A. Adams, 1852: synonym of Montfortula rugosa
  • Emarginula subclathrata Pilsbry, 1890: synonym of Emarginula dilecta A. Adams, 1852
  • Emarginula subemarginata Blainville, 1819: synonym of Montfortia subemarginata
  • Emarginula subrugosa Thiele, 1916: synonym of Hemimarginula subrugosa
  • Emarginula sulcifera A. Adams, 1852: synonym of Montfortulana sulcifera
  • Emarginula tricarinata Pilsbry, 1890: synonym of Montfortista panhi
  • Emarginula tricostata G.B. Sowerby I, 1823: synonym of Hemitoma octoradiata
  • Emarginula vadososinuata Yokoyama, 1922: synonym of Tugalina vadososinuata
  • Emarginula vadum Barnard, 1963: synonym of Emarginula undulata Melvill & Standen, 1903
  • Emarginula variegata A. Adams, 1852: synonym of ''Variemarginula variegata''