Rhipidistia
Rhipidistia, also known as Dipnotetrapodomorpha, is a clade of lobe-finned fishes which includes the tetrapods and lungfishes. Rhipidistia formerly referred to a subgroup of Sarcopterygii consisting of the Porolepiformes and Osteolepiformes, a definition that is now obsolete. However, as cladistic understanding of the vertebrates has improved over the last few decades, a monophyletic Rhipidistia is now understood to include the whole of Tetrapoda and the lungfishes.
Rhipidistia includes Porolepiformes and Dipnoi. Extensive fossilization of lungfishes has contributed to many evolutionary studies of this group. Evolution of autostylic jaw suspension, in which the palatoquadrate bone fuses to the cranium, and the lymph pumping "lymph heart", are unique to this group. Another feature shared by lungfish and tetrapods is the divided atrium, though it evolved convergently.
The precise time at which the choana of tetrapods evolved is debated, with some considering early rhipidistians as the first choanates. The feature is also present in modern lungfish but is probably a case of convergent evolution. The basal stem-lungfish Diabolepis did not possess it. Instead, it had four nostrils like most fish. However, its posterior nares are very close to the lip, meaning a ventral 'displacement' of the posterior nostril can be considered a synapomorphy of the lungfish-tetrapod clade. The complete choana then seems to have developed independently in the two surviving clades.
Etymology
The word "Rhipidistia" is from.The word "Dipnotetrapodomorpha" is from ; from, the combining form of the numeral τέτταρες ; from, the combining form of πούς meaning "foot"; and from, the combining form of μορφή meaning "physical shape".