Cottus (fish)
Cottus is a genus of the mainly freshwater ray-finned fishes belonging to the family Cottidae, the typical sculpins. They are often referred to as the "freshwater sculpins", as they are the principal genus of sculpins to be found in fresh water. They are native to the Palearctic and Nearctic.
They are small fish, mostly less than in length, although a few species can reach twice that size.
Taxonomy
Cottus was first proposed as a genus by Carl Linnaeus in the 10th edition of the Systema Naturae when he described the European bullhead and in 1850 this species was designated as the type species of the genus by the French ichthyologist Charles Frédéric Girard. The 5th edition of the Fishes of the World classifies this genus within the subfamily Cottinae of the family Cottidae. Other authorities have found that the Cottidae, as delimited in the 5th edition of Fishes of the World, is paraphyletic and that the monophyletic grouping is the freshwater sculpins, including the Baikal sculpins, while most of the marine taxa are classified within the family Psychrolutidae.Cottus kazika has been found to be outside of a monophyletic Cottus and has been classified in the monospecific genus Rheopresbe.
The earliest fossil remains of the genus are of †Cottus calcatus from the latest Miocene or early Pliocene of Oregon, US. The species †Cottus cervicornis Storms, 1894 is known from the Early Oligocene of Belgium, but its taxonomic identity is uncertain.
Species
There are currently around 70 recognized species in this genus:- Subgenus Cottus Linnaeus, 1758
- * Cottus aturi Freyhof, Kottelat & Nolte, 2005
- * Cottus cyclophthalmus Sideleva, Kesminas & Zhidkov, 2022
- * Cottus duranii Freyhof, Kottelat & Norte, 2005
- * Cottus dzungaricus Kottelat, 2006
- * Cottus ferrugineus Heckel & Kner, 1857
- * Cottus gobio Linnaeus, 1758
- * Cottus gratzianowi Sideleva, Naseka & Zhidkov, 2015
- * Cottus haemusi Marinov & Dikov, 1986
- * Cottus hispaniolensis Băcescu & Băcescu-Mester, 1964
- * Cottus jaxartensis Berg, 1916
- * Cottus koshewnikowi Gratzianov, 1907
- * Cottus metae Freyhof, Kottelat & Nolte, 2005
- * Cottus microstomus Heckel, 1837
- * Cottus perifretum Freyhof, Kottelat & Nolte,, 2005
- * Cottus petiti Băcescu & Băcescu-Mester, 1964
- * Cottus rhenanus Freyhof, Kottelat & Nolte, 2005
- * Cottus ricei E. W. Nelson, 1876
- * Cottus rondeleti Freyhof, Kottelat & Nolte, 2005
- * Cottus sabaudicus Sideleva, 2009
- * Cottus scaturigo Freyhof, Kottelat & Nolte, 2005
- * Cottus sibiricus Kessler, 1889
- * Cottus spinulosus Kessler, 1872
- * Cottus transsilvaniae Freyhof, Kottelat & Nolte, 2005
- Subgenus Cephalocottus Gratzianov, 1907
- * Cottus amblystomopsis P. Y. Schmidt, 1904
- * Cottus nozawae Snyder, 1911
- Subgenus Cottopsis Girard 1850
- * Cottus aleuticus C. H. Gilbert, 1896
- * Cottus asper J. Richardson, 1836
- * Cottus asperrimus Rutter, 1908
- * Cottus gulosus Girard, 1854
- * Cottus klamathensis C. H. Gilbert, 1898
- * Cottus ohlone Moyle & Campbell, 2022
- * Cottus perplexus C. H. Gilbert & Evermann, 1894
- * Cottus pitensis R. M. Bailey & C. E. Bond, 1963
- * Cottus princeps C. H. Gilbert, 1898
- * Cottus tenuis Evermann & Meek, 1898
- Subgenus Uranidea DeKay, 1842
- * Cottus baileyi C. R. Robins, 1961
- * Cottus bairdii Girard, 1850
- * Cottus bendirei T. H. Bean, 1881
- * Cottus caeruleomentum Kinziger, Raesly & Neely, 2000
- * Cottus carolinae T. N. Gill, 1861
- * Cottus chattahoochee Neely, J. D. Williams & Mayden, 2007
- * Cottus cognatus J. Richardson, 1836
- * Cottus echinatus R. M. Bailey & C. E. Bond, 1963
- * Cottus extensus R. M. Bailey & C. E. Bond, 1963
- * Cottus girardi C. R. Robins, 1961
- * Cottus hubbsi R. M. Bailey & Dimick, 1949
- * Cottus hypselurus C. R. Robins & H. W. Robison, 1985
- * Cottus immaculatus Kinziger & R. M. Wood, 2010
- * Cottus kanawhae C. R. Robins, 2005
- * Cottus paulus J. D. Williams, 2000
- * Cottus rhotheus R. Smith, 1882
- * Cottus specus G. L. Adams & Burr, 2013
- * Cottus tallapoosae Neely, J. D. Williams & Mayden, 2007
- Subgenus Incertae sedis
- * Cottus altaicus Kaschenko, 1899
- * Cottus beldingii C. H. Eigenmann & R. S. Eigenmann, 1891
- * Cottus confusus R. M. Bailey & C. E. Bond, 1963
- * Cottus czerskii L. S. Berg, 1913
- * Cottus dorofeevi Sideleva & Shidkov, 2024
- * Cottus greenei C. H. Gilbert & Culver, 1898
- * Cottus hangiongensis T. Mori, 1930
- * Cottus kolymensis Sideleva & A. Goto, 2012
- * Cottus koreanus R. Fujii, Y. Choi & Yabe, 2005
- * Cottus leiopomus C. H. Gilbert & Evermann, 1894
- * Cottus marginatus T. H. Bean, 1881
- * Cottus nasalis L. S. Berg, 1933
- * Cottus perplexus C. H. Gilbert & Evermann, 1894
- * Cottus pollux Günther, 1873
- * Cottus reinii Hilgendorf, 1879
- * Cottus schitsuumsh M. Lemoine, M. K. Young, McKelvey, L. Eby, Pilgrim & M. K. Schwartz, 2014