Coptodera

Coptodera is a genus of beetles in the family Carabidae, constituted of 105 species distributed across North and South America, Africa, Oceania and eastern Asia.
They are small flattened beetles. The body size is usually under 11 mm and pronotum is generally wider than long. Many of them present patches or spots of different colours on the elytra.
A single fossil species of this genus has been described: Coptodera elektra, from Europe's Eocene, was described in 2015 from a single piece of Baltic amber.
It contains the following species:Coptodera acutipennis Coptodera aeneorufa Bates, 1869 Coptodera aerata Dejean, 1825 Coptodera alluaudi Coptodera alticola Baehr, 2007 Coptodera amoenula Boheman, 1848 Coptodera andrewesi Jedlicka, 1934 Coptodera apicalis Shpeley & Ball, 1993 Coptodera aurata Chevrolat, 1835 Coptodera australis Chaudoir, 1870 Coptodera banjaran Kirschenhofer, 2014 Coptodera banjaranensis Kirschenhofer, 2012 Coptodera basilewskyi Coptodera bastai Baehr, 2014 Coptodera baumi Jedlicka, 1955 Coptodera bifasciata Putzeys, 1845 Coptodera braziliensis Shpeley & Ball, 1993 Coptodera brunnea Shpeley & Ball, 1993 Coptodera catalai Coptodera cechovskyi Kirschenhofer, 2010 Coptodera chalcites Bates, 1869 Coptodera championi Bates, 1883 Coptodera chaudoiri Andrewes, 1919 Coptodera confusa Coptodera congoana Burgeon, 1937 Coptodera congoensis Burgeon, 1937 Coptodera consobrina Coptodera crockerensis Kirschenhofer, 2014 Coptodera cupreotincta Bates, 1869 Coptodera cyanella Bates, 1869 Coptodera dromioides Coptodera elongata Putzeys, 1845 Coptodera eluta Andrewes, 1923 Coptodera emarginata Dejean, 1825 Coptodera equestris Boheman, 1848 Coptodera erwini Shpeley & Ball, 1993 Coptodera esakii Coptodera farai Jedlicka, 1964 Coptodera fasciata Boheman, 1848 Coptodera fasciolata Coptodera festiva Dejean, 1825 Coptodera flavipes Louwerens, 1953 Coptodera flexuosa Schmidt-Goebel, 1846 Coptodera foveolata Shpeley & Ball, 1993 Coptodera fulminans Coptodera grossa Darlington, 1968 Coptodera hieroglyphica Fauvel, 1882 Coptodera hova Alluaud, 1936 Coptodera immaculata Coptodera impicta Chaudoir, 1870 Coptodera indica Coptodera interrupta Schmidt-Goebel, 1846 Coptodera intrusa Hansen, 1968 Coptodera japonica Bates, 1883 Coptodera jimnae Baehr, 2007 Coptodera johorensis Kirschenhofer, 2010 Coptodera kelabitensis Kirschenhofer, 2012 Coptodera lamingtonensis Baehr, 2007 Coptodera laticollis Coptodera legorskyi Kirschenhofer, 2012 Coptodera levrati Coptodera lineata Coptodera lineatocollis Burgeon, 1937 Coptodera lineolata Bates, 1869 Coptodera luzonensis Jedlicka, 1934 Coptodera maculata Dupuis, 1912 Coptodera maculosa Kirschenhofer, 2012 Coptodera marginata Dupuis, 1912 Coptodera mastersii Coptodera maynei Burgeon, 1937 Coptodera megalops Bates, 1869 Coptodera mersingensis Kirschenhofer, 2010 Coptodera monteithi Baehr, 2007 Coptodera montisferrei Baehr, 2007 Coptodera nigrosignata Coptodera nigrostriata Coptodera nigroviridis Shpeley & Ball, 1993 Coptodera nitidipennis Baehr, 2007 Coptodera nitidula Coptodera occulta Hunting & Yang, 2019 Coptodera ocellata Chaudoir, 1870 Coptodera ornatipennis Louwerens, 1962 Coptodera osakana Coptodera ovipennis Louwerens, 1956 Coptodera oxyptera Chaudoir, 1870 Coptodera pakitza Shpeley & Ball, 1993 Coptodera papuella Darlington, 1968 Coptodera perakensis Kirschenhofer, 2012 Coptodera phuongensis Kirschenhofer, 1994 Coptodera picea Dejean, 1826 Coptodera picta Chaudoir, 1870 Coptodera poecila Bates, 1883 Coptodera proksi Jedlicka, 1964 Coptodera relucens Bates, 1869 Coptodera rufescens Buquet, 1835 Coptodera rugiceps Coptodera sahlbergi Chaudoir, 1870 Coptodera sallei Shpeley & Ball, 1993 Coptodera sapaensis Kirschenhofer, 1994 Coptodera schaumii Chaudoir, 1861 Coptodera sexguttata Horvatovich, 1976 Coptodera seyrigi Alluaud, 1936 Coptodera shimianensis Kirschenhofer, 2012 Coptodera sigillata Shpeley & Ball, 1993 Coptodera simillima Baehr, 2007 Coptodera squiresi Coptodera stockwelli Shpeley & Ball, 1993 Coptodera subapicalis Putzeys, 1877 Coptodera subapicaloides Jedlicka, 1956 Coptodera subcostata Louwerens, 1952 Coptodera taiwana Coptodera tetrastigma Chaudoir, 1870 Coptodera teutonica Shpeley & Ball, 1993 Coptodera transversa Coptodera triloba Coptodera tripartita Chaudoir, 1870 Coptodera tripunctata Shpeley & Ball, 1993 Coptodera umbrina Coptodera undulata Perty, 1830 Coptodera uttaranchalensis Kirschenhofer, 2012 Coptodera versicolor Bates, 1869 Coptodera viridana Jedlicka, 1935 Coptodera viridis Shpeley & Ball, 1993 Coptodera wau Darlington, 1968 Coptodera waytkowskii Liebke, 1951 Coptodera xanthopleura Bates, 1891

† Coptodera elektra Gamboa & Ortuño, 2015