Copelatus
Copelatus is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in color, many species of Copelatus possessing visible longitudinal furrows down the dorsal side of the wings of both sexes.
Systematics
The genus Copelatus is divided into several subgenera, some of which are sometimes treated as separate genera. It may be paraphyletic with respect to the smaller Copelatinae genera Lacconectus and Aglymbus. The species include:Species
Copelatus abonnenci Guignot, 1939Copelatus acamas Guignot, 1955Copelatus adelbert Megna, Atthakor, Manaono, Hendrich & Balke, 2017Copelatus advena Sharp, 1882Copelatus aemulus Bilardo & Rocchi, 1995Copelatus aequatorius Régimbart, 1899Copelatus aethiopicus Régimbart, 1906Copelatus agaroensis Balke, 2008Copelatus agrias Guignot, 1954Copelatus alternatus Sharp, 1882Copelatus amaroides Guignot, 1952Copelatus amatolensis Omer-Cooper, 1965Copelatus amazonicus Régimbart, 1889Copelatus ambiguus Bertrand & Legros, 1975Copelatus amphibius Ranarilalatiana & Bergsten, 2019Copelatus anastomosans Guignot, 1952Copelatus andobonicus Guignot, 1960Copelatus andreinii Régimbart, 1905Copelatus andrewesi J. Balfour-Browne, 1939Copelatus angolensis Peschet, 1924Copelatus angouei Bilardo & Rocchi, 2011Copelatus angustus Gschwendtner, 1932Copelatus ankaratra Ranarilalatiana & Bergsten, 2019Copelatus annobomensis J. Balfour-Browne, 1939Copelatus anthracinus Régimbart, 1895Copelatus antoniorum Hájek & Brancucci, 2011- †Copelatus aphroditae Balke, 2003Copelatus apicinotatus Bilardo & Rocchi, 2018Copelatus assalyi Bilardo & Rocchi, 2010Copelatus assimilis Régimbart, 1895Copelatus ateles Guignot, 1955Copelatus atrosulcatus Régimbart, 1906Copelatus babyrousa Hájek, Hendrich & Balke, 2018Copelatus bacchusi Wewalka, 1981Copelatus bacillifer Guignot, 1955Copelatus baculiformis Guignot, 1955Copelatus badeni Sharp, 1882Copelatus bakewelli J. Balfour-Browne, 1939Copelatus baoulicus Bilardo & Pederzani, 1978Copelatus bapounouensis Bilardo & Rocchi, 2019Copelatus baranensis Hájek, Shaverdo, Hendrich & Balke, 2021Copelatus barbouri Young, 1942Copelatus basifasciatus Régimbart, 1895Copelatus basilewskyi Bilardo & Pederzani, 1979Copelatus bechynei Guignot, 1953Copelatus befasicus Guignot, 1956Copelatus bengalensis Guignot, 1955Copelatus bertrandi Nilsson, Bilardo & Rocchi, 1996Copelatus betampona Ranarilalatiana & Bergsten, 2019Copelatus bezdeki Sheth, Ghate & Hájek, 2018Copelatus bibulus Guignot, 1948Copelatus biformis Sharp, 1882Copelatus bimaculatus Copelatus binaghii Bilardo & Pederzani, 1978Copelatus biolleyi Guignot, 1952Copelatus biroi Guignot, 1956Copelatus birougouensis Bilardo & Rocchi, 2018Copelatus biseriatus J. Balfour-Browne, 1939Copelatus biswasi Mukherjee & Sengupta, 1986Copelatus blancasi Guignot, 1958Copelatus blatchleyi Young, 1953 NACopelatus bolivianus Guignot, 1957Copelatus bombycinus Guignot, 1952Copelatus bonvouloiri Sharp, 1882Copelatus bottegoi Régimbart, 1895Copelatus bougainvillensis Hájek, Shaverdo, Hendrich & Balke, 2021Copelatus boukali Hendrich & Balke, 1998Copelatus brancuccii Rocchi, 1979Copelatus brasiliensis Zimmermann, 1921Copelatus brendelli Hájek, Hendrich & Balke, 2018Copelatus brevicornis Copelatus brevistrigatus Guignot, 1959Copelatus brivioi Rocchi, 1976Copelatus bromeliarum Copelatus brullei Aubé, 1838Copelatus brunneus J. Balfour-Browne, 1939Copelatus buqueti Aubé, 1838Copelatus burgeoni Gschwendtner, 1930Copelatus caelatipennis Aubé, 1838Copelatus caelatus Guignot, 1952Copelatus caffer J. Balfour-Browne, 1939Copelatus calaquei Bilardo & Rocchi, 2008Copelatus camerunensis Guignot, 1941Copelatus capensis Sharp, 1882Copelatus carayoni Legros, 1949Copelatus carinatus Sharp, 1882Copelatus celinoides Guignot, 1952Copelatus cessaima Caetano, Bená & Vanin, 2013Copelatus chevrolati Aubé, 1838 NACopelatus chibcha Guignot, 1952Copelatus chinensis Régimbart, 1899Copelatus chipiriricus Guignot, 1957Copelatus chloroticus Régimbart, 1899Copelatus cinnamomeus Régimbart, 1895Copelatus clarki Sharp, 1882Copelatus collarti Gschwendtner, 1932Copelatus compertus Guignot, 1956Copelatus concii Bilardo, 1982Copelatus concolor Sharp, 1882Copelatus concolorans J. Balfour-Browne, 1939Copelatus confinis Bilardo & Rocchi, 1999Copelatus congo Gschwendtner, 1938Copelatus consimilis Bilardo & Rocchi, 2002Copelatus consors Sharp, 1882
- †Copelatus convexus Copelatus cooperae Bilardo & Pederzani, 1972Copelatus cordovai Megna & Epler, 2012Copelatus cordylinoides J. Balfour-Browne, 1938Copelatus coxalis Sharp, 1882Copelatus crassus Régimbart, 1895Copelatus cryptarchoides Régimbart, 1899Copelatus cubaensis Schaeffer, 1908Copelatus curtistriatus Bilardo & Rocchi, 1995Copelatus curvispinis Bilardo & Rocchi, 2008Copelatus cynthiae Bilardo & Rocchi, 2008Copelatus daemeli Sharp, 1882Copelatus danyi Megna & Epler, 2012Copelatus darlingtoni Young, 1942Copelatus davidi Wewalka, 2017Copelatus debilis Sharp, 1882Copelatus deccanensis Sheth, Ghate & Hájek, 2018Copelatus decellei Bilardo, 1982Copelatus decemsulcatus Régimbart, 1895Copelatus deceptor Bilardo & Rocchi, 1995Copelatus dentatipenis Jiang, Hájek & Jia, 2022Copelatus depressus Sharp, 1882Copelatus descarpentriesi Bertrand & Legros, 1975Copelatus diffisus Guignot, 1939Copelatus dimorphus Sharp, 1882Copelatus distinctus Aubé, 1838Copelatus distinguendus Régimbart, 1903Copelatus diversistriatus Jiang, Hájek & Jia, 2022Copelatus djiboutensis Wewalka & Jäch, 2017Copelatus dolosus Guignot, 1956Copelatus doriae Sharp, 1882Copelatus doudouensis Bilardo & Rocchi, 2015Copelatus duodecimstriatus Aubé, 1838Copelatus duponti Aubé, 1838Copelatus efoutensis Bilardo & Rocchi, 1995Copelatus ejactus Omer-Cooper, 1965Copelatus ellai Bilardo & Rocchi, 1995Copelatus elutus Guignot, 1958Copelatus enganensis Guignot, 1940Copelatus epactus Guignot, 1948Copelatus erichsonii Guérin-Méneville, 1847Copelatus esteriensis Bilardo & Pederzani, 1978Copelatus eucritus Guignot, 1952Copelatus evanidus Bilardo & Rocchi, 1995Copelatus exaratus Sharp, 1882Copelatus externus Kirsch, 1873Copelatus falciformis Pederzani & Schizzerotto, 2017Copelatus fasciatus Bilardo & Rocchi, 1995Copelatus fastidiosus Guignot, 1959Copelatus feae Régimbart, 1888Copelatus felicis Hájek, Jiang & Jia, 2022Copelatus fernandensis Guignot, 1952Copelatus ferruginicollis Régimbart, 1895Copelatus ferus Guignot, 1953Copelatus festae Régimbart, 1899Copelatus fidschiensis Zimmermann, 1928Copelatus fijiensis Guignot, 1955Copelatus filiformis Sharp, 1882Copelatus fizpaci Bilardo & Rocchi, 2013Copelatus flavicans Guignot, 1952Copelatus flavidus Régimbart, 1895Copelatus fluviaticus Guignot, 1957Copelatus fontanus J. Balfour-Browne, 1950
- †Copelatus fossilis Říha, 1974Copelatus fractistriatus Bilardo & Rocchi, 1995Copelatus freudei Guignot, 1955Copelatus fryi J. Balfour-Browne, 1939Copelatus fulviceps J. Balfour-Browne, 1938Copelatus fuscipennis Sharp, 1882Copelatus fuscomaculatus Guignot, 1952Copelatus gabonicus Bilardo & Pederzani, 1978Copelatus galapagoensis G.R. Waterhouse, 1845Copelatus garambanus Guignot, 1955Copelatus gardineri Scott, 1912Copelatus geayi Régimbart, 1904Copelatus geniculatus Sharp, 1882Copelatus gentilis Sharp, 1882Copelatus gestroi Copelatus gigas Bilardo & Rocchi, 2019Copelatus glyphicus NACopelatus griffinii Régimbart, 1899Copelatus gschwendtneri Guignot, 1939Copelatus guadelupensis Legros, 1948Copelatus guerini Aubé, 1838Copelatus guineensis Guignot, 1955Copelatus haemorrhoidalis Régimbart, 1883Copelatus hararensis Guignot, 1952Copelatus hardenbergi J. Balfour-Browne, 1950Copelatus hebeter Guignot, 1953Copelatus heterogynus Régimbart, 1899Copelatus hydroporoides Copelatus ibrahimi Zalat, Saleh, Angus & Kaschef, 2000Copelatus ignacioi Hájek, Hendrich & Balke, 2022Copelatus iguelaensis Bilardo & Rocchi, 2002Copelatus ilybioides Régimbart, 1895Copelatus imitator Bilardo & Rocchi, 2002Copelatus inaequalis Sharp, 1882Copelatus incognitus Sharp, 1882Copelatus indicus Sharp, 1882Copelatus inopinatus Bilardo & Rocchi, 1995Copelatus inornatus Sharp, 1882Copelatus insidiosus Bilardo & Rocchi, 1995Copelatus insolitus Chevrolat, 1863Copelatus instabilis Régimbart, 1897Copelatus instriatus Zimmermann, 1921Copelatus insuetus Guignot, 1941Copelatus insulanus Guignot, 1939Copelatus integer Sharp, 1882Copelatus internus Régimbart, 1904Copelatus interstriatus J. Balfour-Browne, 1939Copelatus inuber Guignot, 1952Copelatus ipiformis Régimbart, 1895Copelatus irinus Régimbart, 1899Copelatus irregularis W.J. Macleay, 1871Copelatus ischius Guignot, 1956Copelatus jactator Guignot, 1955Copelatus jaechi Bilardo & Rocchi, 2016Copelatus jamaicensis Young, 1942Copelatus japonicus Sharp, 1884Copelatus jarrigei Legros, 1954Copelatus javanus Régimbart, 1883Copelatus jocosus Guignot, 1955Copelatus johannis Guignot, 1954Copelatus kalaharii Gschwendtner, 1935Copelatus kammuriensis Tamu & Tsukamoto, 1955Copelatus kaszabi Guignot, 1956Copelatus kely Ranarilalatiana & Bergsten, 2019Copelatus kietensis Hájek, Shaverdo, Hendrich & Balke, 2021Copelatus kilimandjarensis Bilardo & Pederzani, 1972Copelatus kindianus Guignot, 1954Copelatus kongouensis Bilardo & Rocchi, 1999Copelatus koreanus T. Mori, 1932Copelatus laccophilinus Sharp, 1882Copelatus laeticulus Sharp, 1882Copelatus laevipennis Hájek, Shaverdo, Hendrich & Balke, 2021Copelatus lamottei Legros, 1954Copelatus lanzai Bilardo & Rocchi, 1995Copelatus laraensis Bilardo & Rocchi, 1995Copelatus lasckonyi Bilardo & Rocchi, 1995Copelatus latens Guignot, 1952Copelatus laticollis Régimbart, 1899Copelatus latifasciatus Bilardo & Rocchi, 1999Copelatus latipes Sharp, 1882Copelatus latus J. Balfour-Browne, 1950Copelatus leonardii Bilardo & Rocchi, 1999Copelatus leonensis Legros, 1954Copelatus lepersonneae Gschwendtner, 1943Copelatus lignosus Guignot, 1955Copelatus lineatipennis Guignot, 1955Copelatus lineatus Copelatus longicornis Sharp, 1882Copelatus lootensi Guignot, 1955Copelatus louayensis Bilardo & Rocchi, 2004Copelatus luctuosus Guignot, 1939Copelatus luridescens Régimbart, 1889Copelatus luteocinctus Guignot, 1955Copelatus luteomaculatus Guignot, 1956Copelatus luzonicus Guignot, 1952Copelatus macellus Guignot, 1950Copelatus madoni Guignot, 1952Copelatus mahajanga Pederzani & Hájek, 2005Copelatus mahleri Holmen & Vazirani, 1990Copelatus makokouensis Bilardo & Rocchi, 1995Copelatus malaisei Guignot, 1954Copelatus malavergnorum Manuel & Ramahandrison, 2022Copelatus mancus Sharp, 1887Copelatus marginatus Sharp, 1882Copelatus marginipennis Copelatus martinbaehri Hendrich, Shaverdo, Hájek & Balke, 2019Copelatus martini Manuel, Deler-Hernández, Megna & Hájek, 2018Copelatus masculinus Régimbart, 1899Copelatus massaicus Guignot, 1941Copelatus mathani Guignot, 1952Copelatus maushomi Sheth, Ghate & Hájek, 2018Copelatus mbokoensis Bilardo & Rocchi, 2006Copelatus melanogrammus Régimbart, 1883Copelatus mendax Bilardo & Rocchi, 2011Copelatus miaowangi Hájek, Jiang & Jia, 2022Copelatus minimus Bilardo & Rocchi, 1995Copelatus minor Bilardo & Pederzani, 1978Copelatus minutissimus J. Balfour-Browne, 1939Copelatus mkambati Bilton & Mlambo, 2022Copelatus mocquerysi Régimbart, 1895Copelatus mohelicus Guignot, 1959Copelatus monticola Guignot, 1951Copelatus montivagus Young, 1942Copelatus mopanshanensis Jiang, Zhao & Hájek, 2022Copelatus mulangensis Bameul, 2003Copelatus mundus Sharp, 1882Copelatus mutabilis Bilardo & Rocchi, 1999Copelatus mvoungensis Bilardo & Rocchi, 2004Copelatus mysorensis Vazirani, 1970Copelatus nakamurai Guéorguiev, 1970Copelatus nangaensis Guignot, 1952Copelatus nanlingensis Hájek, Jiang & Jia, 2022Copelatus nauclerus Guignot, 1939Copelatus neavei J. Balfour-Browne, 1950Copelatus neelumae Vazirani, 1973Copelatus neogestroi Balke, 2008Copelatus neoguineensis Zimmermann, 1919Copelatus nguemai Bilardo & Rocchi, 2011Copelatus nigricans Sharp, 1882Copelatus nigrolineatus Sharp, 1882Copelatus nigropennis Zimmermann, 1927Copelatus nigrostriatus Régimbart, 1895Copelatus nilssoni Wewalka, 1982Copelatus nimbaensis Legros, 1958Copelatus nitens Bilardo & Rocchi, 1999Copelatus nitidus Sharp, 1882Copelatus nodieri Régimbart, 1895Copelatus normalis Erichson, 1847Copelatus notius Omer-Cooper, 1965Copelatus nzei Bilardo & Rocchi, 1999Copelatus oblitus Sharp, 1882Copelatus obscurus Sharp, 1882Copelatus occultus Bilardo & Rocchi, 1995Copelatus ogasawarensis Kamiya, 1932Copelatus onorei Pederzani & Rocchi, 1982Copelatus ornatipennis Zimmermann, 1926Copelatus owas Régimbart, 1895Copelatus pachys Guignot, 1955Copelatus pallidus Régimbart, 1895Copelatus paludorum Guignot, 1952Copelatus pandanorum Scott, 1912Copelatus pantosi Guignot, 1958Copelatus papuensis J. Balfour-Browne, 1939Copelatus parabaptus Guignot, 1955Copelatus parallelipipedus Régimbart, 1895Copelatus parallelus Zimmermann, 1920Copelatus pardii Rocchi, 1990Copelatus parisii Guignot, 1959Copelatus parumstriatus Gschwendtner, 1934Copelatus paryphes Guignot, 1955Copelatus pederzanii Bilardo & Rocchi, 1995Copelatus pereirai Guignot, 1955Copelatus peridinus Guignot, 1955Copelatus pinnifer Guignot, 1952Copelatus piriensis Omer-Cooper, 1965Copelatus platynotus Régimbart, 1895Copelatus polystrigus Sharp, 1882
- †Copelatus ponomarenkoi Říha, 1974Copelatus portior Guignot, 1956Copelatus posticatus Copelatus poungai Bilardo & Rocchi, 2006
- †Copelatus predaveterus K.B. Miller, 2003Copelatus prolixus Sharp, 1882Copelatus prolongatus Sharp, 1882Copelatus propino Guignot, 1960Copelatus propinquus Régimbart, 1895Copelatus proximus Sharp, 1882Copelatus pseudostriatus Ranarilalatiana & Bergsten, 2019Copelatus pulchellus Copelatus pulicarius Régimbart, 1895Copelatus pumilus Régimbart, 1895Copelatus punctatus Bilardo & Rocchi, 1995Copelatus punctulatus Aubé, 1838 NACopelatus puzhelongi Jiang, Hájek & Jia, 2022Copelatus quadrisignatus Régimbart, 1877Copelatus quadristriatus Pederzani & Schizzerotto, 2017Copelatus racenisi Guignot, 1951Copelatus ragazzii Régimbart, 1887Copelatus regimbarti Branden, 1884Copelatus restrictus Sharp, 1882Copelatus rimosus Guignot, 1952Copelatus rivalis Guignot, 1952Copelatus rocchii Bilardo, 1982Copelatus romani Zimmermann, 1924Copelatus rosulae Hájek, Jiang & Jia, 2022Copelatus royi Legros, 1958Copelatus rubiginosus Guignot, 1954Copelatus ruficapillus Régimbart, 1895Copelatus ruteri Legros, 1954Copelatus saegeri Guignot, 1955Copelatus safiotra Ranarilalatiana & Bergsten, 2019Copelatus sahlbergi J. Balfour-Browne, 1939Copelatus sallaei Sharp, 1882Copelatus sanfilippoi Bilardo & Pederzani, 1972Copelatus saverii Pederzani & Schizzerotto, 2017Copelatus scalptus Guignot, 1954Copelatus scaphites Guignot, 1955Copelatus schaefferi Young, 1942Copelatus schereri Wewalka, 1981Copelatus schuhi Hendrich & Balke, 1998Copelatus scutatus Guignot, 1952Copelatus scytalotus Guignot, 1956Copelatus senegalensis Lagar & Hernando, 1991Copelatus sexstriatus Sharp, 1882Copelatus sharpi Branden, 1884Copelatus sibelaemontis Hájek, Hendrich, Hawlitschek & Balke, 2010Copelatus silvestrii Régimbart, 1903Copelatus simoni Régimbart, 1889Copelatus singularis Bilardo & Rocchi, 1995Copelatus sociennus J. Balfour-Browne, 1952Copelatus solitarius Sharp, 1882Copelatus sordidipennis Régimbart, 1895Copelatus speciosus Régimbart, 1892Copelatus spoliatus Guignot, 1955
- †Copelatus stavropolitanus Říha, 1974Copelatus stillicidicola Bilardo & Rocchi, 1995Copelatus striatellus Boheman, 1848Copelatus striaticollis Lucas, 1857Copelatus striatopterus Aubé, 1838Copelatus striatulus Aubé, 1838Copelatus strigipennis Copelatus strigosulus Copelatus strinatii Guignot, 1958Copelatus striolatus Peschet, 1917Copelatus stygis Guignot, 1958Copelatus subdeficiens Régimbart, 1902Copelatus subsimilis Guignot, 1958Copelatus substriatus Kirsch, 1873Copelatus subterraneus Guéorguiev, 1978Copelatus sudrei Bameul, 2003Copelatus sulcatus Sharp, 1882Copelatus sulcipennis Copelatus sumbawensis Régimbart, 1899Copelatus suppar Guignot, 1956Copelatus supplementaris Régimbart, 1895Copelatus sylvaticoides Bilardo & Rocchi, 2013Copelatus sylvaticus Guignot, 1952Copelatus takakurai Satô, 1985Copelatus tanaus Guignot, 1953Copelatus taprobanicus Wewalka & Vazirani, 1985Copelatus tenebrosus Régimbart, 1880Copelatus tengchongensis Hájek, Jiang & Jia, 2022Copelatus teranishii Kamiya, 1938Copelatus terminalis Sharp, 1882Copelatus ternatensis Régimbart, 1899Copelatus terzanii Bilardo & Rocchi, 2016Copelatus thiriouxi Peschet, 1917Copelatus thrasys Guignot, 1952Copelatus tibialis Sharp, 1882Copelatus tinctor Guignot, 1954Copelatus togoensis Régimbart, 1895Copelatus tomokunii Satô, 1985Copelatus tondangoyei Bilardo & Rocchi, 2013Copelatus tostus J. Balfour-Browne, 1950Copelatus trifilis Guignot, 1958Copelatus triglyphus Guignot, 1955Copelatus trilobatus Régimbart, 1895Copelatus tschaga Bilardo & Pederzani, 1972Copelatus tucuchiensis J. Balfour-Browne, 1939Copelatus tulagicus Guignot, 1942Copelatus uludanuensis Hendrich & Balke, 1995Copelatus undecimstriatus Aubé, 1838Copelatus urceolus Hájek, Shaverdo, Hendrich & Balke, 2021Copelatus usagarensis Zimmermann, 1926Copelatus vagatus Guignot, 1952Copelatus vagestriatus Zimmermann, 1919Copelatus vagus Bilardo & Rocchi, 1995Copelatus validus Sharp, 1882Copelatus vancli Hájek, Hendrich & Balke, 2018Copelatus vanninii Bilardo & Rocchi, 1999Copelatus variegatus Régimbart, 1895Copelatus variistriatus Hájek, Shaverdo, Hendrich & Balke, 2021Copelatus venustus Bilardo & Rocchi, 1995Copelatus vigintistriatus Fairmaire, 1869Copelatus vigintisulcatus Régimbart, 1895Copelatus villiersi Guignot, 1950Copelatus virungaensis Bilardo, 1982Copelatus vitraci Legros, 1948Copelatus vivax Guignot, 1953Copelatus vokoka Ranarilalatiana & Bergsten, 2019Copelatus wallacei J. Balfour-Browne, 1939Copelatus waltoni J. Balfour-Browne, 1950Copelatus wayanadensis Manivannan & Madani, 2011Copelatus wewalkai Holmen & Vazirani, 1990Copelatus weymarni J. Balfour-Browne, 1947Copelatus xanthocephalus Régimbart, 1899Copelatus xiniudong Jiang, Hájek & Jia, 2022Copelatus yacumensis Guignot, 1957Copelatus yaguarete Scheers, 2018Copelatus yombensis Bilardo & Rocchi, 2015Copelatus zadiensis Bilardo & Rocchi, 1995Copelatus zanabato Manuel & Ramahandrison, 2022Copelatus zanatanensis Ranarilalatiana & Bergsten, 2019Copelatus zimmermanni Gschwendtner, 1934