Clavatula


Clavatula is a genus of sea snails, marine gastropod mollusks in the family Clavatulidae.

Description

In this genus, the shell is fusiform, with a well-produced spire. The whorls are coronated with tubercles or short spines at the suture;. The aperture is oval. The columellar lip is smooth, arcuate, and callous behind. The outer lip is thin and arcuated, with the anal sinus situated below the sutural coronal. The operculum is semioval with its nucleus about the middle, on the inner side.
The foot is large, short and obtuse behind. The eyes are placed externally near the extremity of the tentacles. The radula contains a small unicuspid central tooth with single sharp laterals.
The species mostly inhabits the west coast of Africa.

Species

Species within the genus Clavatula include:
  • † Clavatula agassizi
  • † Clavatula agatensis Bellardi, 1877 Clavatula ahuiri Cossignani & Ardovini, 2014Clavatula albasulcata R. Aiken, 2021
  • † Clavatula albucianensis Bellardi, 1877
  • † Clavatula anaglypta
  • † Clavatula apenninica Bellardi, 1877
  • † Clavatula apolloniae
  • †Clavatula aralica
  • † Clavatula ariejansseni Harzhauser, Landau & R. Janssen, 2022 Clavatula asamusiensis Nomura & Zinbo, 1940
  • †Clavatula asperulata
  • † Clavatula atatuerki Harzhauser, Landau & R. Janssen, 2022
  • † Clavatula auingeri Finlay, 1927
  • † Clavatula barnabasi Kovács, Leél-Őssy & Vicián, 2023
  • † Clavatula basilica Bellardi, 1877
  • † Clavatula beauforti Oostingh, 1938
  • † Clavatula berauensis Beets, 1941
  • † Clavatula bicarinata Bellardi, 1877 Clavatula bimarginata
  • † Clavatula boreoromana Kautsky, 1925
  • † Clavatula burdigalensis Cossmann, 1895Clavatula caerulea
  • † Clavatula camillae
  • †Clavatula carinifera Grateloup, 1832
  • † Clavatula circumclusa Bellardi, 1877 Clavatula colini Von Maltzan, 1883
  • † Clavatula complanata Bellardi, 1877
  • †Clavatula concatenata Clavatula congoensis Nolf & Verstraeten, 2008
  • † Clavatula consimilis Bellardi, 1877
  • † Clavatula coppii Bellardi, 1877 Clavatula coronata Lamarck, 1801Clavatula cossignanii Ardovini, 2004
  • † Clavatula danuvii Vicián, Z. Kovács & Stein, 2019Clavatula debilis Clavatula decorata Sowerby III, 1916
  • † Clavatula defossa Bellardi, 1877
  • † Clavatula defrancii Bellardi, 1877 Clavatula diadema
  • † Clavatula djocdjocartae
  • † Clavatula dodecapunctata Vera-Peláez & Lozano-Francisco, 2001
  • † Clavatula dujardini Peyrot, 1938
  • † Clavatula eichwaldi Bellardi, 1877
  • † Clavatula excavata Bellardi, 1877 Clavatula filograna Odhner, 1923Clavatula flammulata Knudsen, 1952
  • † Clavatula flexicosta Bellardi, 1877
  • † Clavatula francisci Clavatula gabonensis Melvill, 1923
  • † Clavatula geniculata Bellardi, 1877
  • † Clavatula glaberrima J.P.S. Grateloup, 1832
  • * †Clavatula glaberrima praecedens
  • † Clavatula gothica Bellardi, 1877 Clavatula gracilior Sowerby II, 1870
  • † Clavatula gracilis Erünal-Erentöz, 1958
  • † Clavatula gradata Clavatula hattenbergeri Nolf & Verstraeten, 2008Clavatula helena Bartsch, 1915
  • †Clavatula helwerdae Ceulemans, Van Dingenen & Landau, 2018
  • † Clavatula heros
  • † Clavatula hirmetzli Z. Kovács & Vicián, 2021
  • †Clavatula iberoechinata Vera-Peláez & Lozano-Francisco, 2001 Clavatula imperialis Lamarck, 1816
  • † Clavatula implexa Bellardi, 1877
  • † Clavatula inedita Bellardi, 1877
  • † Clavatula inornata Bellardi, 1877
  • † Clavatula interrupta
  • † Clavatula irisae Harzhauser, Landau & R. Janssen, 2022
  • † Clavatula isseli Bellardi, 1877
  • † Clavatula istvani Kovács, Leél-Őssy & Vicián, 2023
  • † Clavatula jarzynkae Harzhauser, Landau & R. Janssen, 2022
  • † Clavatula juliae
  • † Clavatula justinae Clavatula knudseni Nolf & Verstraeten, 2007Clavatula kraepelini
  • † Clavatula labiolirata Landau, Harzhauser, İslamoğlu & C. M. Silva, 2013
  • † Clavatula laciniata Bellardi, 1877
  • † Clavatula landaui Vera-Peláez & Lozano-Francisco, 2001 Clavatula lelieuri
  • † Clavatula ligeriana Clavatula malala Bozzetti, 2018
  • † Clavatula manzonii Bellardi, 1877
  • † Clavatula mariae Clavatula martensi Von Maltzan, 1883
  • † Clavatula martinelli Vera-Peláez & Lozano-Francisco, 2001 Clavatula matthiasi Nolf, 2008Clavatula milleti Clavatula muricata Clavatula mystica Clavatula nathaliae Nolf, 2006
  • † Clavatula neogradata Glibert, 1954
  • † Clavatula nodosa Bellardi, 1877
  • † Clavatula obtruta
  • † Clavatula olgae
  • † Clavatula oostinghi Beets, 1942
  • † Clavatula orientoromana Báldi, 1960
  • † Clavatula palatina Strausz, 1954
  • † Clavatula pettkoi Biskupič, 2023 Clavatula petzyae Boyer & Ryall, 2006
  • † Clavatula pusilla Bellardi, 1877
  • † Clavatula pyramidum Cuvillier, 1933 Clavatula regia
  • †Clavatula reginae Hoernes & Auinger, 1891Clavatula retusa R.B. Hinds, 1843
  • † Clavatula romana Clavatula rubrifasciata
  • † Clavatula rugata Bellardi, 1877
  • † Clavatula ruida Bellardi, 1877
  • † Clavatula rusticula Clavatula santhai Z. Kovács & Vicián, 2021 †
  • † Clavatula sceauxensis Landau, Van Dingenen & Ceulemans, 2020
  • † Clavatula seminuda Bellardi, 1877
  • † Clavatula seyithasanensis Landau, Harzhauser, İslamoğlu & C. M. Silva, 2013 Clavatula smithi Knudsen, 1952Clavatula solangeae Bozzetti, 2006
  • † Clavatula sophiae
  • † Clavatula sorini Harzhauser, Landau & R. Janssen, 2022
  • † Clavatula sotterii Clavatula strebeli Knudsen, 1952Clavatula taxea
  • † Clavatula theodori Clavatula tripartita
  • † Clavatula turriculata
  • † Clavatula turriculoides Bellardi, 1877
  • † Clavatula unicostata Bellardi, 1877
  • † Clavatula veronicae Clavatula virgineus
  • † Clavatula vitalisi Strausz, 1955 Clavatula xanteni Nolf & Verstraeten, 2006
;Nomen dubium:Clavatula erecta W. H. Turton, 1932
;Species brought into synonymy :Clavatula albicans Hinds, 1843: synonym of Splendrillia albicans Clavatula amabilis Hinds, 1843: synonym of Tritonoturris amabilis Clavatula argillacea Hinds, 1843: synonym of Etrema argillacea Clavatula auriculifera Lamarck, J.B.P.A. de M. 1816: synonym of Clavus canalicularis Clavatula bella Hinds, 1843: synonym of Bellacythara bella Clavatula boreointerrupta Kautsky, 1925: synonym of † Granulatocincta boreointerrupta