Borthwickia
Borthwickia is genus of flowering plants, containing one species, Borthwickia trifoliata from Yunnan, China and Myanmar. The common name in Chinese is 节蒴木. It is a shrub or small tree with evergreen trifoliate leaves, whitish flowers clustered at the tip of the branches, with many stamens, and thin, knobbly, drooping fruits with many small red seeds.
Description
Borthwickia is a fragrant, evergreen shrub or small tree of high. It has square, light green, later pale yellow branchlets, which are initially covered in dense, short, white hairs, which are lost in the older, cylindrical branches. Leaves are arranged with two on opposite sides of the branchlets, and consist of a usually long leaf stalk and three papery leaflets, each on a stalk of about long. The midveins are raised on the upper surface, flat on the lower surface, where they are also covered in dense white hairs. Each leaflet has seven to nine side veins on each side, and reticulate veinlets in between that are visible from both sides. The top leaflet is usually long and about half as wide. The leaflets at the base are slightly smaller and somewhat asymmetric. Flowers are with several in long racemes at the tip of branchlets, and develop from the base to the tip. The common axis is covered in soft short white hairs, and each flower is set on a 1–1½ cm long flower stalk, which is initially in the axil of a 1–1½ cm long narrow bract. The file to eight sepals are whitish in color and fused into a closed tube with short white velvet hair on both surfaces. These split into two lobes when the flower opens, and are quickly shed. The five to eight erect petals are whitish in color and oval to spoon-shaped, 1½–1¾ cm long, thick at base and thin at the upper margin, up to half as long as the calyx tube. Both anthers and ovary are set above sepals and petals on a of about ½ cm long. Sixty to seventy free stamens are long and carry ovate anthers which open with longitudinal slits. Pollen grains are approximately 30 × 22 μm, with three ridges from pole to pole. Nectar is produced by the stalk between the petals and the stamens. The ovary is linear, 1–1½ cm long, towards the tip with four to six grooves and ridges, and four to six compartments, in each of which are two rows of ovules attached to the axis at the centre. The stigma at the top is seated and indistinct. The fruit is on a long stalk above the scars of the stamens, and is a drooping, knobbly, angular cylindrical capsule, of long and in diameter, with a long beak at the tip. When ripe, the walls of the fruits are shed in longitudinal strips starting from the base, showing the axis with eight to twelve rows of large, initially red, later red-brown kidney-shaped seeds. The plant can be in flower from April to June, and ripe fruits may be present in August and September.Borthwickia differs from the other core Brassicales in having opposite leaves and by having four to six fruit compartments.