Aurochs


The aurochs is an extinct species of bovine, considered to be the wild ancestor of modern domestic cattle. With a shoulder height of up to in bulls and in cows, it was one of the largest herbivores in the Holocene; it had massive elongated and broad horns that reached in length.
The aurochs was part of the Pleistocene megafauna. It probably evolved in Asia and migrated west and north during warm interglacial periods. The oldest-known aurochs fossils date to the Middle Pleistocene. The species had an expansive range spanning from Western Europe and North Africa to the Indian subcontinent and East Asia. The distribution of the aurochs progressively contracted during the Holocene due to habitat loss and hunting, with the last known individual dying in the Jaktorów forest in Poland in 1627.
There is a long history of interaction between aurochs and humans, including archaic hominins like Neanderthals. The aurochs is depicted in Paleolithic cave paintings, Neolithic petroglyphs, Ancient Egyptian reliefs and Bronze Age figurines. It symbolised power, sexual potency and prowess in religions of the ancient Near East. Its horns were used in votive offerings, as trophies and drinking horns.
Two aurochs domestication events occurred during the Neolithic Revolution. One gave rise to the domestic taurine cattle in the Fertile Crescent in the Near East that was introduced to Europe via the Balkans and the coast of the Mediterranean Sea. Hybridisation between aurochs and early domestic cattle occurred during the early Holocene. Domestication of the Indian aurochs led to the zebu cattle that hybridised with early taurine cattle in the Near East about 4,000 years ago. Some modern cattle breeds exhibit features reminiscent of the aurochs, such as the dark colour and light eel stripe along the back of bulls, the lighter colour of cows, or an aurochs-like horn shape.

Etymology

Both "aur" and "ur" are Germanic or Celtic words meaning "wild ox".
In Old High German, this word was compounded with ohso to ūrohso, which became the early modern Aurochs. The Latin word "urus" was used for wild ox from the Gallic Wars onwards.
The use of the plural form aurochsen in English is a direct parallel of the German plural Ochsen and recreates the same distinction by analogy as English singular ox and plural oxen, although aurochs may stand for both the singular and the plural term; both are attested.

Taxonomy and evolution

The scientific name Bos taurus was introduced by Carl Linnaeus in 1758 for feral cattle in Poland.
The scientific name Bos primigenius was proposed for the aurochs by Ludwig Heinrich Bojanus who described the skeletal differences between the aurochs and domestic cattle in 1825, published in 1827. The name Bos namadicus was used by Hugh Falconer in 1859 for cattle fossils found in Nerbudda deposits.
Bos primigenius mauritanicus was coined by Philippe Thomas in 1881 who described fossils found in deposits near Oued Seguen west of Constantine, Algeria.
In 2003, the International Commission on Zoological Nomenclature placed Bos primigenius on the Official List of Specific Names in Zoology and thereby recognized the validity of this name for a wild species.

Subspecies

Three aurochs subspecies have traditionally been recognised to have existed in historical times:
Chinese geneticists published mitochondrial DNA evidence supporting that Eurasian aurochs populations from northern China were genetically isolated for large stretches of the Pleistocene, and as a result distinctive enough to be considered a separate subspecies, the East Asian aurochs, even if this was not morphologically distinct.
At least two dwarf subspecies of aurochs developed on Mediterranean islands as a result of sea level changes during the Pleistocene:
using fossils of 16 Bovidae species indicate that the Bovini tribe evolved about. The Bos and Bison genetic lineages are estimated to have genetically diverged from the Bovini about. The following cladogram shows the phylogenetic relationships of the aurochs based on analysis of nuclear and mitochondrial genomes in the Bovini tribe:
The cold Pliocene climate caused an extension of open grassland, which enabled the evolution of large grazers. The origin of the aurochs is unclear, with authors suggesting either an African or Asian origin for the species. Bos acutifrons is considered to be a possible ancestor of the aurochs, of which a fossil skull was excavated in the Sivalik Hills in India that dates to the Early Pleistocene about.
An aurochs skull excavated in Tunisia's Kef Governorate from early Middle Pleistocene strata dating about is the oldest well-dated fossil specimen to date. The authors of the study proposed that Bos might have evolved in Africa and migrated to Eurasia during the Middle Pleistocene. Middle Pleistocene aurochs fossils were also excavated in a Saharan erg in the Hoggar Mountains.
Fossils of the Indian subspecies were excavated in alluvial deposits in South India dating to the Middle Pleistocene. Remains of aurochs are common in Late Pleistocene sites across the Indian subcontinent.
The earliest fossils in Europe date to the Middle Pleistocene. One site widely historically suggested to represent the first appearance of aurochs in Europe was the Notarchirico site in southern Italy, dating around 600,000 years ago, however a 2024 re-examination of the site found that presence of aurochs at the locality was unsupported, with the oldest records of aurochs now placed at the Ponte Molle site in central Italy, dating to around 550–450,000 years ago. Aurochs were present in Britain by Marine Isotope Stage 11 ~400,000 years ago.
The earliest remains aurochs in East Asia are uncertain, but may date to the late Middle Pleistocene, probably no earlier than 200,000 years ago.
Late Pleistocene aurochs fossils were found in Affad 23 in Sudan dating to 50,000 years ago when the climate in this region was more humid than during the African humid period.
Following the most recent deglaciation, the range of the aurochs expanded into Denmark and southern Sweden at the beginning of the Holocene, around 12–11,000 years ago.

Description

According to a 16th-century description by Sigismund von Herberstein, the aurochs was pitch-black with a grey streak along the back; his wood carving made in 1556 was based on a culled aurochs, which he had received in Mazovia. In 1827, Charles Hamilton Smith published an image of an aurochs that was based on an oil painting that he had purchased from a merchant in Augsburg, which is thought to have been made in the early 16th century. This painting is thought to have shown an aurochs, although some authors suggested it may have shown a hybrid between an aurochs and domestic cattle, or a Polish steer. Contemporary reconstructions of the aurochs are based on skeletons and the information derived from contemporaneous artistic depictions and historic descriptions of the animal.

Coat colour

Remains of aurochs hair were not known until the early 1980s. Depictions show that the North African aurochs may have had a light saddle marking on its back. Calves were probably born with a chestnut colour, and young bulls changed to black with a white eel stripe running down the spine, while cows retained a reddish-brown colour. Both sexes had a light-coloured muzzle, but evidence for variation in coat colour does not exist. Egyptian grave paintings show cattle with a reddish-brown coat colour in both sexes, with a light saddle, but the horn shape of these suggest that they may depict domesticated cattle.
Many primitive cattle breeds, particularly those from Southern Europe, display similar coat colours to the aurochs, including the black colour in bulls with a light eel stripe, a pale mouth, and similar sexual dimorphism in colour. A feature often attributed to the aurochs is blond forehead hairs. According to historical descriptions of the aurochs, it had long and curly forehead hair, but none mentions a certain colour. Although the colour is present in a variety of primitive cattle breeds, it is probably a discolouration that appeared after domestication.

Body shape

The proportions and body shape of the aurochs were strikingly different from many modern cattle breeds. For example, the legs were considerably longer and more slender, resulting in a shoulder height that nearly equalled the trunk length. The skull, carrying the large horns, was substantially larger and more elongated than in most cattle breeds. As in other wild bovines, the body shape of the aurochs was athletic, and especially in bulls, showed a strongly expressed neck and shoulder musculature. Therefore, the fore hand was larger than the rear, similar to the wisent, but unlike many domesticated cattle. Even in carrying cows, the udder was small and hardly visible from the side; this feature is equal to that of other wild bovines.

Size

The aurochs was one of the largest herbivores in Holocene Europe. The size of an aurochs appears to have varied by region, with larger specimens in northern Europe than farther south. Aurochs in Denmark and Germany ranged in height at the shoulders between in bulls and in cows, while aurochs bulls in Hungary reached.
The African aurochs was similar in size to the European aurochs in the Pleistocene, but declined in size during the transition to the Holocene; it may have also varied in size geographically.
The body mass of aurochs appears to have shown some variability. Some individuals reached around, whereas those from the late Middle Pleistocene are estimated to have weighed up to. The aurochs exhibited considerable sexual dimorphism in the size of males and females.