Andricus


Andricus is a genus of oak gall wasps in the family Cynipidae.

Life cycle

As in all Hymenoptera, sex-determination in species of the genus Andricus is governed by haplodiploidy: males develop from unfertilized eggs and are haploid, and females develop from fertilized eggs and are diploid.
Many species in the genus alternate between an asexual generation and a sexual generation.

Species

The taxonomy of the genus Andricus is uncertain, with many of the approximately 375 species considered dubious. The following species are currently recognised in the genus Andricus: Andricus aciculatus Beutenmuller, 1909Andricus ahmeti Melika, Mutun & Dinç, 2014Andricus albipes Hartig, 1840Andricus albobalani Weld, 1926Andricus alniensis Folliot, 1964Andricus amblycerus Andricus amenti Giraud, 1859Andricus analis Andricus anatolicus Melika, Mutun & Dinç, 2014Andricus archiboldi Melika & Abrahamson, 2021Andricus aries Andricus arslani Azmaz & Katılmış, 2022Andricus assarehi Melika & Sadeghi, 2008Andricus atkinsonae Melika, Stone, Sadeghi & Zargaran, 2008Andricus bakrachus Melika, Mutun & Dinç, 2014Andricus balanaspis Andricus balanella Andricus barriosi Medianero & Nieves-Aldrey, 2019Andricus biconicus Weld, 1959Andricus bimaculatus Andricus brachycentrus Andricus breviramuli Pujade-Villar, 2014Andricus brunneus Fullaway, 1911 — clustered gall waspAndricus bulgaricus Vasileva-Symnalieva, 1978Andricus burgundus Giraud, 1859Andricus burnetti Dailey & Sprenger, 1983Andricus caliciformis Andricus californicus Ashmead, 1885Andricus callidoma Andricus capillatus Andricus caputmedusae Andricus catalinensis Melika, Nicholls & Stone, 2021Andricus catilla Andricus cecconii Kieffer, 1901Andricus chapmanii Melika & Abrahamson, 2021Andricus chico Weld, 1957Andricus chinquapin — small oak spindle gall waspAndricus chiricahuensis Melika, Nicholls & Stone, 2021Andricus chodjaii Melika, 2008 Andricus chrysolepidicola Ashmead, 1896 — irregular spindle gall waspAndricus cinnamomeus Ashmead, 1887Andricus clementinae Andricus coconinoensis Melika, Nicholls & Stone, 2021Andricus columbiensis Melika, Nicholls & Stone, 2021Andricus comata Weld, 1956Andricus confusus Lobato-Vila & Pujade-Villar, 2019Andricus confertus McCracken & Egbert, 1922 — convoluted gall wasp Andricus conglomeratus Andricus conificus Andricus cooki Melika, Nicholls & Stone, 2021Andricus coortus Weld, 1947 — club gall waspAndricus coquilletti - little oak apple gall waspAndricus coriariformis Melika, Challis & Stone, 2008Andricus coriarius Andricus coronatus Andricus coronus Beutenmueller, 1907Andricus corruptrix Andricus costaricensis Pujade-Villar & Melika, 2018Andricus costatus Weld, 1944Andricus crassicornis Andricus crispator Tschek, 1871Andricus cryptobius Wachtl, 1880Andricus csokai Melika & Tavakoli, 2008Andricus curtisii Andricus curvator Andricus cydoniae Giraud, 1859Andricus deciduatus Weld, 1926Andricus dentimitratus Andricus dimorphus — clustered midrib gall waspAndricus discalis Andricus exclusus Andricus fidelensis Kieffer, 1901Andricus fitzpatricki Melika & Abrahamson, 2021Andricus flavicornis Schenck, 1863Andricus flavus Pujade-Villar, Wang, Guo & Chen, 2014Andricus floridus Tavares, 1918Andricus foecundatrix Andricus flavohirtus Beutenmüller, 1913Andricus foliaformis Weld, 1927Andricus formosalis Weld, 1947Andricus formosanus Tang & Melika, 2009Andricus forni Pujade-Villar & Nicholls, 2020Andricus frondeum Andricus fulviventris Schenck, 1863Andricus furnaceus Kinsey, 1920Andricus fuscicornis Hartig, 1841Andricus galeatus Andricus gallaecus Tavares, 1916Andricus gallaetinctoriae Andricus gallaeurnaeformis Andricus gemmeus Andricus gemmicola Kieffer, 1901Andricus geniculatus Andricus giardina Stefani, 1898Andricus glandulae Andricus glutinosus Andricus gracilicornis Andricus grossulariae Andricus hartigi Andricus hastata Andricus hedwigia Andricus highlandensis Melika, Nicholls & Stone, 2021Andricus hispanicus Andricus howertoni Bassett, 1890Andricus hungaricus Andricus hyalinus Hartig, 1841Andricus hystrix Kieffer, 1897Andricus impropius Ballido & Pujade-Villar, 2003Andricus incertus Bassett, 1900Andricus indistinctus Bassett, 1890Andricus infectorius Andricus inflator Hartig, 1840Andricus insanus ]Andricus istvani Melika, 2008Andricus kashiwaphilus Abe, 1998Andricus kollari Andricus korlevici Kieffer, 1902Andricus laevigatus Schenck, 1863Andricus lasius - hairy gall waspAndricus lateralis Andricus legitimus Wiebes-Rijks, 1980Andricus libani Melika, Challis & Stone, 2008Andricus lignicolus Andricus longipennis Andricus lucidus Andricus luisieri Tavares, 1914Andricus luteicornis Kieffer, 1899Andricus malpighii Andricus mammadovi Azmaz & Katilmis, 2021Andricus mamillaformis Andricus marmoreus Kinsey, 1920Andricus maxwelli Bassett, 1890Andricus mayri Andricus megatruncicolus Melika, 2008Andricus melikai Pujade-Villar & Kwast, 2002Andricus mellificus Melika, Nicholls & Stone, 2021Andricus mendocinensis Weld, 1957Andricus menkei Melika & Abrahamson, 2021Andricus miriami Shachar, 2015Andricus mitratus Andricus mogollonensis Melika, Nicholls & Stone, 2021Andricus montezumus Beutenmuller, 1913Andricus moniliatus Hartig, 1840Andricus moreae Andricus morula Shachar, Inbar & Dorchin, 2017Andricus mukaigawae Andricus multiplicatus Andricus murtfeldtae Ashmead, 1896Andricus nichollsi Melika & Stone, 2021Andricus niger Andricus nigricens Andricus nobrei Tavares, 1901Andricus nodifex Kieffer, 1900Andricus notholithocarpi Melika, Nicholls & Stone, 2018Andricus occultatus Andricus octosporifex Andricus opertusfimbriate gall waspAndricus ostrea Andricus pallidicornis Andricus pallipes Andricus panteli Andricus paradoxus Andricus partali Pujade-Villar, Cuesta-Porta & Hanson, 2022Andricus pedicellatus Kinsey, 1922 — hair stalk gall waspAndricus peredurus Andricus perfulvum Andricus perlentus Andricus petioli Hartig, 1843Andricus pictus Andricus pilula Andricus pilularis Weld, 1952Andricus pisiformis Beutenmueller, 1911Andricus polycerus Andricus prescotti Weld, 1952Andricus projectus Weld, 1952Andricus pseudoaries Melika, Stone & Sadegh, 2008Andricus pseudocurvator Tang & Melika, 2011Andricus pseudoinflator Tavares, 1901Andricus pujadevillari Melika, Stone, Sadeghi, Atkinson & Zargaran, 2008Andricus quadrilineatus Andricus quercuscalicisknopper gallAndricus quercuscalifornicusCalifornia gall waspAndricus quercuscorticis Andricus quercusfoliatus — leafy oak gall waspAndricus quercusformosus Andricus quercusfrondosus Andricus quercusinferus Andricus quercuslaurinus Melika and Pujade-Villar, 2009Andricus quercuspetioli Andricus quercuspetiolicola Bassett, 1863 — oak petiole gall waspAndricus quercuspyramidalis Andricus quercusradicis Andricus quercusramuli Andricus quercussingularis Bassett, 1863 — small oak apple gall waspAndricus quercustozae Andricus quercusstrobilanuslobed oak gall waspAndricus quercusutriculus Andricus reticulatus Bassett, 1890Andricus rhizoxenus Andricus rhyzomae Andricus robustus Weld, 1926Andricus rotundula Andricus rubripes Andricus ruficornis Andricus rufipes Andricus rufiventris Schenck, 1863Andricus rufus Andricus rugatus Weld, 1926Andricus ruginosus Bassett, 1890Andricus sadeghii Melika, Stone, Atkinson & Aligolizade, 2008 Andricus schencki Dalla Torre & Kieffer, 1910 Andricus schickae Nicholls, Melika & Stone, 2021Andricus schoenroggei Melika & Stone, 2008Andricus schroeckingeri Wachtl, 1876Andricus scutella Weld, 1930Andricus seckendorffi Andricus seminationis Andricus serotinus Andricus serricornis Andricus sessilum Weld, 1926Andricus shuhuti Melika, Mutun & Dinç, 2013Andricus sieboldi Andricus singularis Mayr, 1870Andricus solitarius Andricus songshui Tang & Melika, 2011Andricus sphaericus Pujade-Villar, 2016Andricus spicatus Andricus splendens Weld, 1919Andricus stefanii Andricus stellaris Weld, 1926 — sunburst gall waspAndricus stellatus Melika & Tavakoli, 2008Andricus stellulus — stellar gall waspAndricus sternlichti Bellido, Pujade-Villar & Melika, 2003Andricus stonei Melika, Tavakoli & Sadeghi, 2014Andricus stramineus Weld, 1944Andricus stropus Ashmead, 1887Andricus strues Andricus subterranea Andricus sufflator Mayr, 1882Andricus sulfureus Weld, 1926Andricus superfetationis Andricus surculi Andricus tavaresi Kieffer, 1904Andricus testaceipes Andricus testaceus Andricus texanus Beutenmüller, 1909Andricus theophrasteus Andricus tomentosus Andricus torreyaensis Melika & Abrahamson, 2021Andricus toumeyi Weld, 1926Andricus trotteri Kieffer, 1898Andricus truncicola Andricus tubalis Weld, 1926Andricus tubularius Weld, 1926Andricus tumefaciens Pujade-Villar & Paretas-Martínez, 2014Andricus turcicus Melika, Mutun & Dinç, 2014Andricus turionum Andricus vacciniifoliaeoak apple waspAndricus villosulus Andricus verensis Weld, 1955Andricus vindobonensis Müllner, 1901Andricus weldi Andricus wheeleri Beutenmueller, 1907Andricus williami Melika, Nicholls & Stone, 2021Andricus wiltzae Fullaway, 1911Andricus zappellai Kieffer, 1901

Previous species

These species were previously grouped in Andricus.
Currently in Druon:Druon flocculentum LyonDruon fullawayi Beutenmüller, 1914Druon ignotum BassettDruon linaria KinseyDruon pattoni Druon protagion KinseyDruon quercusflocci Druon quercuslanigerum Ashmead
Currently in Feron:Feron albicomus Feron amphorus Feron apiarium Feron atrimentum Kinsey, 1922) — striped volcano gall waspFeron bakkeri Feron caepula Feron californicum Feron clarkei Feron crystallinum — crystalline gall waspFeron cylindratum Feron discularis Feron gigas —saucer gall waspFeron kingi — red cone gall waspFeron parmula — disc gall waspFeron pattersonae Fullaway, 1911 — plate gall waspFeron tecturnarum Kinsey, 1920Feron tubifaciens Weld, 1926