Amylocorticiales


Amylocorticiales is an order of fungi in the class Agaricomycetes. The order was circumscribed in 2010 to contain mostly resupinate forms that have been referred to genera Anomoporia, Amyloathelia, Amylocorticiellum, Amylocorticium, Amyloxenasma, Anomoloma, Athelopsis, Ceraceomyces, Hypochniciellum, Leptosporomyces and Serpulomyces and the anomalous species, Athelia rolfsii, now classified in its own genus, Agroathelia.

Phylogenetics

The order contains one of three assemblages of basal lineages of Agaricomycetidae that contain corticioid fungi; the others are the Jaapiales and the Atheliales. Although several molecular studies have confirmed the placement of the Amylocorticiales in the Agaricomycetidae, its relationship with other higher-level taxa is not known precisely. Depending on the loci used for analysis, the order is supported as sister group to the Agaricales, or as sister to a clade that contains the Agaricales, Boletales and Atheliales.

Description

Species in the Amylocorticiales form fruit bodies that are effused, effused-reflexed to almost pileate, or stipitate. They have smooth hymenophores that can be either merulioid, irpicoid or poroid. The hyphal system is monomitic and all hyphae are nodose-septate. The cystidia are rare and when present, are tube-like and often nodose-septate. The basidia are mostly terminal but in one genus borne laterally on horizontal hyphae, invariably producing four spores. The basidiospores are smooth, thin-walled or thick-walled, ellipsoid, cylindrical or allantoid, in most species amyloid. Taxa that do not exhibit amyloid reactions include Agroathelia, Ceraceomyces, Podoserpula, Serpulomyces and Leptosporomyces septentrionalis. Species are saprobic on decaying wood or as plant parasites. Amylocorticiales typically cause brown rot or white rot. Agroathelia is a major plant pathogen of field crops

Genera

image:Podoserpula pusio.jpg |thumb|right|Podoserpula pusio is an exception to the typically resupinate fruit body form characteristic of the Amylocorticiales.