Amorphophallus
Amorphophallus is a large genus of some 200 tropical and subtropical tuberous herbaceous plants from the arum family, native to Asia, Africa, Australia and various oceanic islands. The genus includes the Titan arum of Indonesia, which has the largest inflorescence in the genus, and is known as the 'corpse flower' for the pungent odour it produces during its flowering period, which can take up to seven years of growth before it occurs.
A few species are edible as "famine foods" after careful preparation to remove irritating chemicals.
History
The oldest systematic record of the plants was in 1692, when Van Rheede tot Drakenstein published descriptions of two plants. The name "Amorphophallus" was first mentioned in 1834 by the Dutch botanist Blume from Ancient Greek αμορφος amorphos "without form, misshapen" and φαλλος phallos "penis", referring to the shape of the prominent spadix. Between 1876 and 1911, Adolf Engler merged a number of other genera into Amorphophallus, with a final monograph published in 1911.Distribution
These are typical lowland plants, growing in the tropical and subtropical zones of the paleotropics, from West Africa through the Pacific Islands. None of them are found in the Americas, although a remarkably similar but not closely related genus, Dracontium, has evolved there. Most species are endemic. They grow preferentially on disturbed grounds, such as secondary forests.Description
Image:Amorphophallus_sp._fruit.jpg|thumb|150px|Amorphophallus infructescence with berry-like fruitsThese small to massive plants grow from a subterranean tuber. Amorphophallus tubers vary greatly from species to species, from the quite uniformly globose tuber of A. konjac to the elongated tubers of A. longituberosus and A. macrorhizus to the bizarre clustered rootstock of A. coaetaneus. The weight of these tubers range from as little as in Amorphophallus pusillus of Vietnam to as much as for Amorphophallus titanum. From the top of this tuber a single leaf, which can be several metres across in larger species, is produced atop a trunk-like petiole followed, on maturity, by a single inflorescence. This leaf consists of a vertical leaf stalk and a horizontal blade, which may consist of a number of small leaflets. The leaf lasts one growing season. The peduncle can be long or short.
As is typical of the Arum family, these species develop an inflorescence consisting of an elongate or ovate spathe which usually envelops the spadix. The spathe can have different colours, but mostly brownish-purple or whitish-green. On the inside, they contain ridges or warts, functioning as insect traps.
The plants are monoecious. The spadix has tiny flowers: female flowers, no more than a pistil, at the bottom, then male flowers, each with one stamen, and then a blank sterile area. This last part, called 'the appendix', consists of sterile flowers, called staminodes, and can be especially large. The flowers do not have corollas.
Mature female flowers are usually receptive for only one day. In many species, the inflorescence emits a scent of decaying flesh in order to attract insects, though a number of species give off a pleasant odour. Through a number of ingenious insect traps, pollinating insects that entered a spathe when female flowers were receptive remain inside the spathe for about one day while male flowers mature and release pollen. Pollen falls on these insects, and they carry pollen as they exit the spathe and can pollinate female flowers in another spathe. Amorphophallus species are used as food plants by the larvae of some Lepidoptera species including Palpifer sexnotatus and Palpifer sordida.
Pollinated flowers usually each develop into a globose berry, a fruit. The berries are red, orange-red, white, white and yellow, or blue, depending on the species.
Notable species
The species Amorphophallus titanum, 'corpse flower' or titan arum, has the world's largest unbranched inflorescence, with a height of up to and a width of. After an over -tall flower opened at Chicago Botanic Gardens on 29 September 2015, thousands lined up to see and smell it. The floriculturalist described it as smelling "like roadkill, a barnyard, a dirty diaper, very strong, a little bit of mothball smell too". Native to the Indonesian rainforest, it takes about 10 years to blossom. Dubbed "Alice", its bloom was broadcast via live webcam. It is one of two plants at the Chicago Botanic Gardens, which kept open until 2 am on September 30 to accommodate visitors.A runner-up is Amorphophallus gigas, which is taller but has a somewhat smaller inflorescence.
Amorphophallus konjac tubers are used to make konjac, a Japanese thickening agent and edible jelly containing glucomannan.
Some species are called voodoo-lily, as are some species of Typhonium.