Tropical rainforest
Tropical rainforests are dense and warm rainforests with high rainfall typically found between 10° north and south of the Equator. They are a subset of the tropical forest biome that occurs roughly within the 28° latitudes. Tropical rainforests are a type of tropical moist broadleaf forest, that includes the more extensive seasonal tropical forests. True rainforests usually occur in tropical rainforest climates where no dry season occurs; all months have an average precipitation of at least. Seasonal tropical forests with tropical monsoon or savanna climates are sometimes included in the broader definition.
Tropical rainforests ecosystems are distinguished by their consistent, high temperatures, exceeding monthly, and substantial annual rainfall. The abundant rainfall results in nutrient-poor, leached soils, which profoundly affect the flora and fauna adapted to these conditions. These rainforests are renowned for their significant biodiversity. They are home to 40–75% of all species globally, including half of the world's animal and plant species, and two-thirds of all flowering plant species. Their dense insect population and variety of trees and higher plants are notable. Described as the "world's largest pharmacy", over a quarter of natural medicines have been discovered in them. However, tropical rainforests are threatened by human activities, such as logging and agricultural expansion, leading to habitat fragmentation and loss.
The structure of a tropical rainforest is stratified into layers, each hosting unique ecosystems. These include the emergent layer with towering trees, the densely populated canopy layer, the understory layer rich in wildlife, and the forest floor, which is sparse due to low light penetration. The soil is characteristically nutrient-poor and acidic. Tropical rainforests have a long history of ecological succession, influenced by natural events and human activities. They are crucial for global ecological functions, including carbon sequestration and climate regulation. Many indigenous peoples around the world have inhabited rainforests for millennia, relying on them for sustenance and shelter, but face challenges from modern economic activities.
Conservation efforts are diverse, focusing on both preservation and sustainable management. International policies, such as the Reducing Emissions from Deforestation and Forest Degradation programs, aim to curb deforestation and forest degradation. Despite these efforts, tropical rainforests continue to face significant threats from deforestation and climate change, highlighting the ongoing challenge of balancing conservation with human development needs.
Overview
Tropical rainforests are hot and wet. Mean monthly temperatures exceed during all months of the year. Average annual rainfall is no less than and can exceed although it typically lies between and. This high level of precipitation often results in poor soils due to leaching of soluble nutrients in the ground.Tropical rainforests exhibit high levels of biodiversity. Around 40% to 75% of all biotic species are indigenous to the rainforests. Rainforests are home to half of all the living animal and plant species on the planet. Two-thirds of all flowering plants can be found in rainforests. A single hectare of rainforest may contain 42,000 different species of insect, up to 807 trees of 313 species and 1,500 species of higher plants. Tropical rainforests have been called the "world's largest pharmacy", because over one quarter of natural medicines have been discovered within them. It is likely that there may be many millions of species of plants, insects and microorganisms still undiscovered in tropical rainforests.
Tropical rainforests are among the most threatened ecosystems globally due to large-scale fragmentation as a result of human activity. Habitat fragmentation caused by geological processes such as volcanism and climate change occurred in the past, and have been identified as important drivers of speciation. However, fast human driven habitat destruction is suspected to be one of the major causes of species extinction. Tropical rain forests have been subjected to heavy logging and agricultural clearance throughout the 20th century, and the area covered by rainforests around the world is rapidly shrinking.
History
Tropical rainforests have existed on earth for hundreds of millions of years. Most tropical rainforests today are on fragments of the Mesozoic era supercontinent of Gondwana. The breakup of Gondwana left tropical rainforests located in five major regions of the world: tropical America, Africa, Southeast Asia, Madagascar, and New Guinea, with smaller outliers in Australia. However, the specifics of the origin of rainforests remain uncertain due to an incomplete fossil record.Other types of tropical forest
Several biomes may appear similar to, or merge via ecotones with, tropical rainforest:File:Hana Highway Overlook.jpg|thumb|Hawaiian tropical rainforest seen from the Hana Highway
;Moist seasonal tropical forest:
File:Rain Forest Daintree Australia.jpg|thumb|Daintree rainforest in Queensland is a seasonal tropical forest.
Moist seasonal tropical forests receive high overall rainfall with a warm summer wet season and a cooler winter dry season. These forests usually fall under tropical monsoon or tropical savanna climates. Some trees in these forests drop some or all of their leaves during the winter dry season, thus they are sometimes called "tropical mixed forest". They are found in parts of South America, in Central America and around the Caribbean, in coastal West Africa, parts of the Indian subcontinent, and across much of Indochina.
;Montane rainforests:
These are found in cooler-climate mountainous areas, becoming known as cloud forests at higher elevations. Depending on latitude, the lower limit of montane rainforests on large mountains is generally between 1500 and 2500m while the upper limit is usually from 2400 to 3300m.
;Flooded rainforests:
Tropical freshwater swamp forests, or "flooded forests", are found in Amazon basin and elsewhere.
Forest structure
Rainforests are divided into different strata, or layers, with vegetation organized into a vertical pattern from the top of the soil to the canopy. Each layer is a unique biotic community containing different plants and animals adapted for life in that particular strata. Only the emergent layer is unique to tropical rainforests, while the others are also found in temperate rainforests.Forest floor
The forest floor, the bottom-most layer, receives only 2% of the sunlight. Only plants adapted to low light can grow in this region. Away from riverbanks, swamps and clearings, where dense undergrowth is found, the forest floor is relatively clear of vegetation because of the low sunlight penetration. This more open quality permits the easy movement of larger animals such as: ungulates like the okapi, tapir, Sumatran rhinoceros, and apes like the western lowland gorilla, as well as many species of reptiles, amphibians, and insects. The forest floor also contains decaying plant and animal matter, which disappears quickly, because the warm, humid conditions promote rapid decay. Many forms of fungi growing here help decay the animal and plant waste.Understory layer
The understory layer lies between the canopy and the forest floor. The understory is home to a number of birds, small mammals, insects, reptiles, and predators. Examples include leopard, poison dart frogs, ring-tailed coati, boa constrictor, and many species of Coleoptera. The vegetation at this layer generally consists of shade-tolerant shrubs, herbs, small trees, and large woody vines which climb into the trees to capture sunlight. Only about 5% of sunlight breaches the canopy to arrive at the understory causing true understory plants to seldom grow to 3 m. As an adaptation to these low light levels, understory plants have often evolved much larger leaves. Many seedlings that will grow to the canopy level are in the understory.Canopy layer
The canopy is the primary layer of the forest, forming a roof over the two remaining layers. It contains the majority of the largest trees, typically 30–45 m in height. Tall, broad-leaved evergreen trees are the dominant plants. The densest areas of biodiversity are found in the forest canopy, as it often supports a rich flora of epiphytes, including orchids, bromeliads, mosses and lichens. These epiphytic plants attach to trunks and branches and obtain water and minerals from rain and debris that collects on the supporting plants. The fauna is similar to that found in the emergent layer, but more diverse. It is suggested that the total arthropod species richness of the tropical canopy might be as high as 20 million. Other species inhabiting this layer include many avian species such as the yellow-casqued wattled hornbill, collared sunbird, grey parrot, keel-billed toucan, scarlet macaw as well as other animals like the spider monkey, African giant swallowtail, three-toed sloth, kinkajou, and tamandua.Emergent layer
The emergent layer contains a small number of very large trees, called emergents, which grow above the general canopy, reaching heights of 45–55m, although on occasion a few species will grow to 70–80m tall. Some examples of emergents include: Hydrochorea elegans, Dipteryx panamensis, Hieronyma alchorneoides, Hymenolobium mesoamericanum, Lecythis ampla and Terminalia oblonga. These trees need to be able to withstand the hot temperatures and strong winds that occur above the canopy in some areas. Several unique faunal species inhabit this layer such as the crowned eagle, the king colobus, and the large flying fox.However, stratification is not always clear. Rainforests are dynamic and many changes affect the structure of the forest. Emergent or canopy trees collapse, for example, causing gaps to form. Openings in the forest canopy are widely recognized as important for the establishment and growth of rainforest trees. It is estimated that perhaps 75% of the tree species at La Selva Biological Station, Costa Rica are dependent on canopy opening for seed germination or for growth beyond sapling size, for example.