Extinction
Extinction is the termination of a species via the death of its last member. A taxon may become functionally extinct before the death of its last member if it loses the capacity to reproduce and recover. As a species' potential range may be very large, determining this moment is difficult, and is usually done retrospectively. This difficulty leads to phenomena such as Lazarus taxa, where a species presumed extinct abruptly "reappears" after a period of apparent absence.
Over five billion species are estimated to have died out. It is estimated that there are currently around 8.7 million species of eukaryotes globally, possibly many times more if prokaryotes are included. Notable extinct animal species include non-avian dinosaurs, saber-toothed cats, and mammoths. Through evolution, species arise through the process of speciation. Species become extinct when they are no longer able to survive in changing conditions or against superior competition. The relationship between animals and their ecological niches has been firmly established. A typical species becomes extinct within 10 million years of its first appearance, although some species, called living fossils, survive with little to no morphological change for hundreds of millions of years, though this claim has been disputed.
Mass extinctions are relatively rare events; however, isolated extinctions of species and clades are quite common, and are a natural part of the evolutionary process. Only recently have extinctions begun to be recorded, and there is an ongoing mass extinction event caused by human activity. Most species that become extinct are never scientifically documented. Some scientists estimate that up to half of presently existing plant and animal species may become extinct by 2100. A 2018 report indicated that the phylogenetic diversity of 300 mammalian species erased during the human era since the Late Pleistocene would require 5 to 7 million years to recover.
According to the 2019 Global Assessment Report on Biodiversity and Ecosystem Services by IPBES, the biomass of wild mammals has fallen by 82%, natural ecosystems have lost about half their area and a million species are at risk of extinction—all largely as a result of human actions. Twenty-five percent of plant and animal species are threatened with extinction. In a subsequent report, IPBES listed unsustainable fishing, hunting and logging as being some of the primary drivers of the global extinction crisis. In June 2019, one million species of plants and animals were at risk of extinction. At least 571 plant species have been lost since 1750. The main cause of the extinctions is the destruction of natural habitats by human activities, such as cutting down forests and converting land into fields for farming.
A dagger symbol placed next to the name of a species or other taxon normally indicates its status as extinct.
Definition
A species is extinct when the last existing member dies. Extinction therefore becomes a certainty when there are no surviving individuals that can reproduce and create a new generation. A species may become functionally extinct when only a handful of individuals survive, which cannot reproduce due to poor health, age, sparse distribution over a large range, a lack of individuals of both sexes, or other reasons.Pinpointing the extinction of a species requires a clear definition of that species. If it is to be declared extinct, the species in question must be uniquely distinguishable from any ancestor or daughter species, and from any other closely related species. Extinction of a species plays a key role in the punctuated equilibrium hypothesis of Stephen Jay Gould and Niles Eldredge.
File:Various dinosaurs.png|thumb|left|Skeleton of various extinct dinosaurs; some other dinosaur lineages still flourish in the form of birds
In ecology, extinction is sometimes used informally to refer to local extinction, in which a species ceases to exist in the chosen area of study, despite still existing elsewhere. Local extinctions may be made good by the reintroduction of individuals of that species taken from other locations; wolf reintroduction is an example of this. Species that are not globally extinct are termed extant. Those species that are extant, yet are threatened with extinction, are referred to as threatened or endangered species.
File:Edwards' Dodo.jpg|thumb|The dodo of Mauritius, shown here in a 1626 illustration by Roelant Savery, is an often-cited example of modern extinction.
Currently, an important aspect of extinction is human attempts to preserve critically endangered species. These are reflected by the creation of the conservation status "extinct in the wild". Species listed under this status by the International Union for Conservation of Nature are not known to have any living specimens in the wild and are maintained only in zoos or other artificial environments. Some of these species are functionally extinct, as they are no longer part of their natural habitat and it is unlikely the species will ever be restored to the wild. When possible, modern zoological institutions try to maintain a viable population for species preservation and possible future reintroduction to the wild, through use of carefully planned breeding programs.
The extinction of one species' wild population can have knock-on effects, causing further extinctions. These are also called "chains of extinction". This is especially common with extinction of keystone species.
A 2018 study indicated that the sixth mass extinction started in the Late Pleistocene could take up to 5 to 7 million years to restore mammal diversity to what it was before the human era.
Pseudoextinction
Extinction of a parent species where daughter species or subspecies are still extant is called pseudoextinction or phyletic extinction. Effectively, the old taxon vanishes, transformed into a successor, or split into more than one.Pseudoextinction is difficult to demonstrate unless one has a strong chain of evidence linking a living species to members of a pre-existing species. For example, it is sometimes claimed that the extinct Hyracotherium, which was an early horse that shares a common ancestor with the modern horse, is pseudoextinct, rather than extinct, because there are several extant species of Equus, including zebra and donkey; however, as fossil species typically leave no genetic material behind, one cannot say whether Hyracotherium evolved into more modern horse species or merely evolved from a common ancestor with modern horses. Pseudoextinction is much easier to demonstrate for larger taxonomic groups.
Lazarus taxa
A Lazarus taxon or Lazarus species refers to instances where a species or taxon was thought to be extinct, but was later rediscovered. It can also refer to instances where large gaps in the fossil record of a taxon result in fossils reappearing much later, although the taxon may have ultimately become extinct at a later point.The coelacanth, a fish related to lungfish and tetrapods, is an example of a Lazarus taxon that was known only from the fossil record and was considered to have been extinct since the end of the Cretaceous Period. In 1938, however, a living specimen was found off the Chalumna River on the east coast of South Africa. Calliostoma bullatum, a species of deepwater sea snail originally described from fossils in 1844 proved to be a Lazarus species when extant individuals were described in 2019.
Attenborough's long-beaked echidna is an example of a Lazarus species from Papua New Guinea that had last been sighted in 1962 and believed to be possibly extinct, until it was recorded again in November 2023.
Some species thought to be extinct have had ongoing speculation that they may still exist, and in the event of rediscovery would be considered Lazarus species. Examples include the thylacine, or Tasmanian tiger, the last known example of which died in Hobart Zoo in Tasmania in 1936; the Japanese wolf, last sighted over 100 years ago; the American ivory-billed woodpecker, with the last universally accepted sighting in 1944; and the slender-billed curlew, not seen since 2007.
Causes
As long as species have been evolving, species have been going extinct. It is estimated that over 99.9% of all species that ever lived are extinct. The average lifespan of a species is 1–10 million years, although this varies widely between taxa.A variety of causes can contribute directly or indirectly to the extinction of a species or group of species. "Just as each species is unique", write Beverly and Stephen C. Stearns, "so is each extinction ... the causes for each are varied—some subtle and complex, others obvious and simple". Most simply, any species that cannot survive and reproduce in its environment and cannot move to a new environment where it can do so, dies out and becomes extinct. Extinction of a species may come suddenly when an otherwise healthy species is wiped out completely, as when toxic pollution renders its entire habitat unliveable; or may occur gradually over thousands or millions of years, such as when a species gradually loses out in competition for food to better adapted competitors. Extinction may occur a long time after the events that set it in motion, a phenomenon known as extinction debt.
Assessing the relative importance of genetic factors compared to environmental ones as the causes of extinction has been compared to the debate on nature and nurture. The question of whether more extinctions in the fossil record have been caused by evolution or by competition or by predation or by disease or by catastrophe is a subject of discussion; Mark Newman, the author of Modeling Extinction, argues for a mathematical model that falls in all positions. By contrast, conservation biology uses the extinction vortex model to classify extinctions by cause. When concerns about human extinction have been raised, for example in Sir Martin Rees' 2003 book Our Final Hour, those concerns lie with the effects of climate change or technological disaster.
Human-driven extinction started as humans migrated out of Africa more than 60,000 years ago. Currently, environmental groups and some governments are concerned with the extinction of species caused by humanity, and they try to prevent further extinctions through a variety of conservation programs. Humans can cause extinction of a species through overharvesting, pollution, habitat destruction, introduction of invasive species, overhunting, and other influences. Explosive, unsustainable human population growth and increasing per capita consumption are essential drivers of the extinction crisis. According to the International Union for Conservation of Nature, 784 extinctions have been recorded since the year 1500, the arbitrary date selected to define "recent" extinctions, up to the year 2004; with many more likely to have gone unnoticed. Several species have also been listed as extinct since 2004.