Forest raven
The forest raven, also commonly known as the Tasmanian raven, is a passerine bird in the family Corvidae native to Tasmania and parts of southern Victoria, such as Wilsons Promontory and Portland. Populations are also found in parts of New South Wales, including Dorrigo and Armidale. Measuring in length, it has all-black plumage, beak and legs. As with the other two species of raven in Australia, its black feathers have grey bases. Adults have white irises; younger birds have dark brown and then hazel irises with an inner blue rim. New South Wales populations are recognised as a separate subspecies C. tasmanicus boreus, but appear to be nested within the Tasmanian subspecies genetically.
The forest raven lives in a wide variety of habitats in Tasmania but is restricted to more closed forest on mainland Australia. Breeding takes place in spring and summer, occurring later in Tasmania than in New South Wales. The nest is a bowl-shaped structure of sticks sited high in a tree. An omnivorous and opportunistic feeder, the forest raven eats a wide variety of plant and animal material, as well as food waste from urban areas and roadkill. It has been blamed for killing lambs and poultry and raiding orchards in Tasmania, and is unprotected under Tasmanian legislation. The forest raven is sedentary, with pairs generally bonding for life and establishing permanent territories.
Taxonomy and naming
described the "South-Seas raven" in 1781, with loose throat feathers and found in "the Friendly Isles" in the South Seas, but did not give a binomial name. Although "the Friendly Isles" refers to Tonga, the specimen resembles what is now known as the forest raven and was collected by ships' surgeon William Anderson on the third voyage of James Cook in January 1777. Of the species, he had written, "Crows, nearly the same as ours in England". Tasked as the expedition's naturalist, Anderson collected many bird specimens but had died of tuberculosis in 1778 before the return home. Many collection localities were incorrect, and notes were lost or pieced together many years later. German naturalist Johann Friedrich Gmelin gave the species the name Corvus australis in the 13th edition of Systema naturae in 1788.Since Australia was settled by Europeans, all species of crows and ravens have been colloquially known as crows by the general population and are difficult to distinguish. In his 1865 Handbook to the Birds of Australia John Gould noted a single species of corvid in Australia, Corvus australis, which he called the white-eyed crow. He used Gmelin's 1788 name, which took precedence by virtue of its age over Vigors and Horsfield's description. In 1912 Scottish naturalist William Robert Ogilvie-Grant clarified the species as C. coronoides and C. cecilae. Subsequently, French-American ornithologist Charles Vaurie acted as First Revisor under Article 24 of the International Code of Zoological Nomenclature Code and discarded C. australis as a junior homonymin 1788 Gmelin had used the same binomial name to describe the black nunbirdto preserve the stability of the name. This has been followed by later authors.
Gregory Mathews described the forest raven as a distinct subspecies of the Australian raven in 1912, its species name derived from Tasmania, the type locality. Ian Rowley raised the forest raven to species rank in 1970, noting there were no intermediate forms between it and the little raven and that it was clearly larger with a much more massive bill. He described a second subspecies the same year, observing that C. tasmanicus from Tasmania and southern Victoria has a very short tail compared with individuals from the northern New South Wales population. The term "crow" is colloquially applied to any or all species of Australian corvid. In 1970 Rowley gave the species name "forest raven", which was later designated the official name by the International Ornithologists' Union.
Preliminary genetic analysis of the genus using mitochondrial DNA showed the three raven species to belong to one lineage and the two crows to another, and that the two lineages are not closely related. The genetic separation between species is small and there was a suggestion the forest raven may be conspecificity with the Australian raven. Subsequent multigene analysis using nuclear DNA by Jønsson and colleagues in 2012 clarified that the forest and little raven are each other's closest relative. The northern subspecies boreus turned out to be nested in the Tasmanian tasmanicus, indicating the populations separated very recently. It is still recognised as a distinct subspecies by the International Ornithological Committee.
Ian Rowley proposed that the common ancestor of the five species diverged into a tropical crow and temperate raven sometime after entering Australia from the north. The raven diverged into the ancestor of the forest and little ravens in the east and Australian raven in the west. As the climate was cooler and drier, the aridity of central Australia split them entirely as the habitat between became inhospitable. Furthermore, the eastern diverged into nomadic little ravens and, in forested refuges, forest ravens. As the climate eventually became warmer, the western ravens spread eastwards and outcompeted forest ravens on mainland Australia, as evidenced by the forest ravens' being found only in closed forest refuges on the mainland but in a wider variety of habitats in Tasmania.
Description
The largest of the Australian corvids, the adult forest raven is long with a wingspan of and weighing approximately. There is no seasonal variation in plumage, which is entirely glossy black with a blue or green sheen visible on the upperparts. The wings are long and broad, with the largest of its ten primary feathers almost reaching the end of the tail when at rest. The tail is rounded or wedge-shaped. It is quite short in Tasmanian populations but longer in northern New South Wales. The beak is a similar shape to that of the little raven, though more massive and heavy-set. The upper mandible, including the nares and nasal groove, is covered with bristles. The mouth and tongue are black, as are the powerful legs and feet. The tibia is fully feathered and the tarsus is long.Sexes have identical plumage; the male is generally larger, but there is considerable overlap in size between individuals. The forest raven can be distinguished from the two species of crow occurring in Australia by the grey base of the feathers, which is white in the latter species. The demarcation between pale and black regions on the feather is gradual in the ravens and sharply delineated in the crows. Feather bases are not normally visible when observing birds in the field, but can sometimes be seen on a windy day if the feathers are ruffled. The three species of raven are more heavily set with a broader chest than the two crow species, with the forest raven the stockiest of all. Relative size is useful only when two species can be seen side by side, as the overlap in size is large and the difference in size small. In Tasmania, the forest raven could be confused with the black currawong, though the latter species has more slender wings with white markings, a longer tail and a very different call.
Juveniles have a shorter, shallower bill, which is dark grey with some pink at the base. The gape is pink. The plumage is softer and fluffier and often has a brown tint. It generally lacks the glossy sheen of adult birds, though a blue-purple sheen can be seen sometimes on mantle and shoulders plumage. Birds between one and two years old closely resemble adults but retain juvenile feathers on wings and tail and have smaller bills. Birds between two and three years have adult plumage but lack the adult eye colour. Eye colour varies with age: nestlings up to four months old have blue-grey eyes, juveniles aged from four to fourteen months have brown eyes, and immature birds have hazel eyes with blue eyerings around the pupil until age two years and ten months.
Vocalization
The call is considered the most reliable means of identification in areas where the forest raven's range overlaps with other corvids. It is a deep and husky "korr-korr-korr-korr" with a similarly drawn out last note to the Australian raven. It can also utter a barking alarm call. The calls of juveniles have a higher pitch than those of adult birds. Mated pairs greet each other with a specific return-home call; a long extended descending call, and characteristic flapping flight with reduced-amplitude wingbeats.Distribution and habitat
The only member of the corvid family that has a permanent population in Tasmania, the forest raven is the most widely distributed bird species in the state. There are three populations in southern Victoria: from the vicinity of Lakes Entrance west across Gippsland to Wilsons Promontory, the Otway Ranges from west of Torquay to Port Campbell, and lastly in the Grampians and Millicent Plain extending into south-east South Australia. Isolated records suggest the latter two populations may actually be continuous. There are two disjunct populations in northern New South Wales. A coastal population is found from Tea Gardens north to Yuraygir National Park, while a more montane population is found along the Great Dividing Range and New England Tableland from Gloucester Tops in the south to Tenterfield in the north. The gap between the two populations is around, shrinking to at Dorrigo.The forest raven inhabits a wide range of habitat within Tasmania such as woods, open interrupted forest, mountains, coastal areas, farmland and town and city fringes. A survey of Mount Wellington found it to be one of the few birds that remained in open and marshland habitat at higher elevations over the winter. Additionally, research within Tasmania found that ravens were thirty percent more likely to be observed in farmland habitat than in non-agricultural forested or urban areas. On mainland Australia it appears to be more confined to forests: wet and dry sclerophyll forest and cool temperate rainforest, as well as pine plantations in Victoria. Populations in Victoria and New South Wales are possibly expanding, with the species more evident in towns such as Forster-Tuncurry and Port Macquarie, and along segments of the Oxley Highway between Wauchope and Walcha, and Thunderbolts Way between Gloucester and Nowendoc, most likely due to roadkill from increased vehicular traffic. It is unclear whether records since the 1970s in areas where the forest raven was unknown are the result of range expansion or improved field observations and identification.
Forest ravens fly from Tasmania and the mainland to islands well offshore in Bass Strait and may even traverse the strait entirely. First recorded on King Island in Bass Strait in 1979, the forest raven has become more numerous and flocks of several hundred birds were recorded by 1997. The island was previously inhabited by little ravens.