Atlantic cod
The Atlantic cod is a fish of the family Gadidae, widely consumed by humans. It is also commercially known as cod or codling.
In the western Atlantic Ocean, cod has a distribution north of Cape Hatteras, North Carolina, and around both coasts of Greenland and the Labrador Sea; in the eastern Atlantic, it is found from the Bay of Biscay north to the Arctic Ocean, including the Baltic Sea, the North Sea, Sea of the Hebrides, areas around Iceland and the Barents Sea.
Atlantic cod can live for up to 25 years and typically grow up to, but individuals in excess of and have been caught. They will attain sexual maturity between ages two and eight with this varying between populations and has varied over time.
Colouring is brown or green, with spots on the dorsal side, shading to silver ventrally. A stripe along its lateral line is clearly visible. Its habitat ranges from the coastal shoreline down to along the continental shelf.
Atlantic cod is one of the most heavily fished species. Atlantic cod was fished for a thousand years by north European fishers who followed it across the North Atlantic Ocean to North America. It supported the US and Canadian fishing economy until 1992, when the Canadian Government implemented a ban on fishing cod.
Several cod stocks collapsed in the 1990s and have failed to fully recover even with the cessation of fishing. This absence of the apex predator has led to a trophic cascade in many areas. Many other cod stocks remain at risk. The Atlantic cod is labelled vulnerable on the IUCN Red List of Threatened Species, per a 1996 assessment that the IUCN notes needs updating. A 2013 assessment covering only Europe shows the Atlantic cod has rebounded in Europe, and it has been relabelled least concern.
Dry cod may be prepared as unsalted stockfish, and as cured salt cod or clipfish.
Taxonomy
The Atlantic cod is one of three cod species in the genus Gadus along with Pacific cod and Greenland cod. A variety of fish species are colloquially known as cod, but they are not all classified within the Gadus, though some are in the Atlantic cod family, Gadidae.Behaviour
Shoaling
Atlantic cod are a shoaling species and move in large, size-structured aggregations. Larger fish act as scouts and lead the shoal's direction, particularly during post spawning migrations inshore for feeding. Cod actively feed during migration and changes in shoal structure occur when food is encountered. Shoals are generally thought to be relatively leaderless, with all fish having equal status and an equal distribution of resources and benefits. However, some studies suggest that leading fish gain certain feeding benefits. One study of a migrating Atlantic cod shoal showed significant variability in feeding habits based on size and position in the shoal. Larger scouts consumed a more variable, higher quantity of food, while trailing fish had less variable diets and consumed less food. Fish distribution throughout the shoal seems to be dictated by fish size, and ultimately, the smaller lagging fish likely benefit from shoaling because they are more successful in feeding in the shoal than they would be if migrating individually, due to social facilitation.Predation
Atlantic cod are apex predators in the Baltic and adults are generally free from the concerns of predation. Juvenile cod, however, may serve as prey for adult cod, which sometimes practice cannibalism. Juvenile cod make substrate decisions based on risk of predation. Substrates refer to different feeding and swimming environments. Without apparent risk of predation, juvenile cod demonstrated a preference for finer-grained substrates such as sand and gravel-pebble. However, in the presence of a predator, they preferred to seek safety in the space available between stones of a cobble substrate. Selection of cobble significantly reduces the risk of predation. Without access to cobble, the juvenile cod simply tries to escape a predator by fleeing.Additionally, juvenile Atlantic cod vary their behaviour according to the foraging behaviour of predators. In the vicinity of a passive predator, cod behaviour changes very little. The juveniles prefer finer-grained substrates and otherwise avoid the safer kelp, steering clear of the predator. In contrast, in the presence of an actively foraging predator, juveniles are highly avoidant and hide in cobble or in kelp if cobble is unavailable.
Heavy fishing of cod in the 1990s and the collapse of American and Canadian cod stocks resulted in trophic cascades. As cod are apex predators, overfishing them removed a significant predatory pressure on other Atlantic fish and crustacean species. Population-limiting effects on several species including American lobsters, crabs, and shrimp from cod predation have decreased significantly, and the abundance of these species and their increasing range serve as evidence of the Atlantic cod's role as a major predator rather than prey.
Swimming
Atlantic cod have been recorded to swim at speeds of a minimum of and a maximum of with a mean swimming speed of. In one hour, cod have been recorded to cover a mean range of. Swimming speed was higher during the day than at night. This is reflected in the fact that cod more actively search for food during the day. Cod likely modify their activity pattern according to the length of daylight, thus activity varies with time of year.Response to changing temperatures
Swimming and physiological behaviours change in response to fluctuations in water temperature. Respirometry experiments show that heart rates of Atlantic cod change drastically with changes in temperature of only a few degrees. A rise in water temperature causes marked increases in cod swimming activity. Cod typically avoid new temperature conditions, and the temperatures can dictate where they are distributed in water. They prefer to be deeper, in colder water layers during the day, and in shallower, warmer water layers at night. These fine-tuned behavioural changes to water temperature are driven by an effort to maintain homeostasis to preserve energy. This is demonstrated by the fact that a decrease of only caused a highly costly increase in metabolic rate of 15–30%.Feeding and diet
The diet of the Atlantic cod consists of fish such as herring, capelin, and sand eels, as well as squid, mussels, clams, tunicates, comb jellies, brittle stars, sand dollars, sea cucumbers, crustaceans, and polychaetes. Stomach sampling studies have discovered that small Atlantic cod feed primarily on crustaceans, while large Atlantic cod feed primarily on fish. In certain regions, the main food source is decapods with fish as a complementary food item in the diet. Wild Atlantic cod throughout the North Sea depend, to a large extent, on commercial fish species also used in fisheries, such as Atlantic mackerel, haddock, whiting, Atlantic herring, European plaice, and common sole, making fishery manipulation of cod significantly easier. Ultimately, food selection by cod is affected by the food item size relative to their own size. However, providing for size, cod do exhibit food preference and are not simply driven by availability.Atlantic cod practice some cannibalism. In the southern North Sea, 1–2% of stomach contents for cod larger than consisted of juvenile cod. In the northern North Sea, cannibalism was higher, at 10%. Other reports of cannibalism have estimated as high as 56% of the diet consists of juvenile cod.
When hatched, cod larvae are altricial, entirely dependent on a yolk sac for sustenance until mouth opening at ~24 degree days. The stomach generally develops at around 240 degree days. Before this point the intestine is the main point of food digestion using pancreatic enzymes such as trypsin.
Reproduction
Atlantic cod will attain sexual maturity between ages two and eight with this varying between different populations and has also varied over time with a population. Their gonads take several months to develop and most populations will spawn from January to May. For many populations, the spawning grounds are located in a different area from the feeding grounds so require the fish to migrate in order to spawn. On the spawning area, males and females will form large schools. Based on behavioral observations of cod, the cod mating system has been likened to a lekking system, which is characterized by males aggregating and establishing dominance hierarchies, at which point females may visit and choose a spawning partner based on status and sexual characteristics. Evidence suggests male sound production and other sexually selected characteristics allow female cod to actively choose a spawning partner. Males also exhibit aggressive interactions for access to females.Atlantic cod are batch spawners, in which females will spawn approximately 5–20 batches of eggs over a period of time with 2–4 days between the release of each batch. Each female will spawn between 2 hundred thousand and 15 million eggs, with larger females spawning more eggs. Females release gametes in a ventral mount, and males then fertilize the released eggs. The eggs and newly hatched larvae float freely in the water and will drift with the current, with some populations relying upon the current to transport the larvae to nursery areas.