Anoual Formation
The Anoual Formation is a geological formation in the High Atlas of Morocco. It is early Bathonian in age. It consists of two members. The lower member is several hundred metres thick, and consists largely of mudstone with lens beds of cross bedded sandstone, with thin intercalations of limestone that was deposited in a freshwater continental setting, likely lacustrine or palustrine, with small marine influences. The upper member is several tens of metres thick and consists of limestone deposited in a shallow marine setting. The formation is fossiliferous, with several of the limestone intercalations yielding a diverse fauna, including amphibians, reptiles, dinosaurs and mammals.
Paleobiota
Brachiopoda
Crustacea
Mollusca
Fish
Abundant osteichthyan ichthyoliths are known. An isolated spine displaying characters seen in extant cypriniform or siluriform fishes has been recovered, alternatively can belong to a member of Myriacanthidae.| Genus | Species | Location | Stratigraphic Position | Material | Notes | Images |
| Arganodus | Indeterminate | Guelb el Ahmar | Upper Member | Isolated tooth plates | Lungfish | |
| Lepidotes/''Scheenstia'' | Indeterminate | Guelb el Ahmar | Upper Member | Isolated teeth and scales | Lepidotidae bony fish | |
| cf. Ionoscopiformes | Indeterminate | Guelb el Ahmar | Upper Member | Rhomboidal scales | Tentatively assigned to Ophiopsidae | |
| Mawsoniidae | Indeterminate | Guelb el Ahmar | Upper Member | Skull bones including a parasphenoid | Coelacanth | |
| Osteoglossiformes | Indeterminate | Guelb el Ahmar | Upper Member | Squamules | The oldest record of the group |